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中亚热带天然阔叶用材林目标树经营研究
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摘要
我国中亚热带天然阔叶林分布广泛,植物区系组成丰富,具有重要的生态、经济、社会价值。本研究以福建永安天然米槠林为研究对象,探讨目标树经营在天然米槠林应用中的相关问题,如天然米槠林林分特征、目标树经营依据、目标直径、目标树密度、目标树经营对单木、林分的影响等,以期为我国中亚热带天然阔叶林可持续经营提供参考。研究取得以下结论:
     (1)天然米槠林的复杂林分特征是经营的基础,根据天然米槠林划分的林层为基础,研究了天然米槠林物种组成、物种多样性及重要值、直径结构、树高结构、蓄积组成、空间结构等林分特征,为目标树经营提供依据和参考。林分特征主要内容有:天然米槠林明显分化为三个层次,第Ⅰ林层树高≥18m,第Ⅱ林层树高在10.1-16.9m之间,第Ⅲ林层树高≤10m;米槠、木荷、新木姜子、杜英、猴欢喜和丝栗栲是林分该群落的优势种;天然米槠林的第Ⅰ亚层、第Ⅱ亚层、第Ⅲ亚层直径结构使用正态分布、Weibull分布拟合效果最好,全林直径结构Gamma分布、负指数分布拟合最好;全林及主要树种混交频度最大值均在中度混交或强度混交,天然米槠林林分大小比数均值为0.48,天然米槠林林分角尺度均值为0.75。
     (2)选择少量优秀个体林木作为经营主体而把大部分林木留给自然调控的目标树经营是具有生态合理性和经济有效性。通过分析发现天然米槠林第I林层林木株数只占林分总株数的15%,却提供了林分70%以上的活立木蓄积量,这种垂直结构分布特征表明能够提供森林经营物质收获的主体部分在为数不多的大径级林木。采用“平均生长量+标准偏差”的方法和数据拟合胸径不同径阶生长的函数模型,结果证明林分的生长过程主要是由生长较快的优势木执行完成的,即胸径在生长旺盛时期的大径级林木具有更快的生长趋势。
     (3)通过定量方法计算,研究了天然米槠林当前阶段和收获阶段目标树密度问题。基于Voronoi图计算目标树竞争指数和优势度,通过筛选得到当前阶段天然米槠林平均单位面积目标树密度为105株/hm2,确定当前阶段天然米槠林合理目标树密度为100~110株/hm2。基于目标树生长发育至目标直径所需单元营养面积获得收获阶段天然米槠林平均单株营养面积、平均单元营养面积、平均目标树密度为143.05m2、155.49m2、68株/hm2。利用逐步回归方法模拟目标树冠幅,获得天然米槠林目标树达到收获阶段时目标树密度为68~74株/hm2。最后分析认为后两种方法获得的收获阶段天然米槠林目标树密度是合理的,确定天然米槠林收获阶段目标树合适密度为63~73株/hm2左右。
     (4)经过分析择伐后目标树与非目标树生长量、生长率,发现目标树生长量明显高于非目标树生长量,并且明显促进了目标树生长率的增加,证明干扰树择伐后可以明显促进目标树的生长。胸径、蓄积连年生长量和平均生长量排序为目标树T>非目标树NTI>非目标树NT>非目标树NTII>非目标树NTIII,目标树T与其它非目标树生长差异显著。目标树与非目标树断面积生长率、蓄积生长率大小排序为非目标树NT>目标树T>非目标树NTI,择伐2年后目标树断面积、蓄积生长率最大,与择伐3.5年后目标树断面积、蓄积生长率差异不显著,但与择伐1年后目标树断面积、蓄积生长率差异显著。
     (5)分析2007-2011年不同经营模式断面积、蓄积生长率,结果表明全林及各林层断面积生长率排序为目标树经营>传统经营>对照。对照全林断面积生长率与传统经营和目标树经营全林断面积生长率差异显著,但后两种经营模式全林断面积生长率差异不显著;全林及各林层蓄积生长率大小顺序为目标树经营>传统经营>对照,对照全林蓄积生长率与传统经营和目标树经营全林蓄积生长率差异显著,但后两种经营模式全林生长率差异不显著。
     (6)分析不同经营模式前后重要值的变化情况,结果表明三种模式经营前后米槠、木荷仍然为最主要的优势树种。对照样带米槠、木荷、尖叶水丝梨、少叶黄桤、蓝果树重要值均小幅度下降,新木姜子、猴欢喜、光叶山矾重要值增加幅度较大;传统经营样带米槠、木荷、新木姜子、山黄皮、尖叶水丝梨重要值下降幅度较大;目标树经营样带米槠、木荷、浙江润楠、拟赤杨、薯豆、厚壳桂重要值都呈现下降趋势,新木姜子、华杜英、枫香、虎皮楠、树参、兰果树重要值呈增加趋势。
It has widely distribution and aboundance of resource of natural broad-leavead forest inour mid-subtropical zone and the forest resources have important ecological, economic, socialvalue.The natural broad-leavead Castanopsis carlesii in Yong’an is taken as study objectiveaim to resolve realated problems of target management. Such as charictaristics of naturalbroad-leavead Castanopsis carlesii, basis of target management, target tree diameter, target treedensity, target tree management effect on single and stand. The study hopes to provide areference for sustaninable manangement of broad-levead forest in our mid-subtropical zone.The study made the following conclusions:
     (1) Basis on the different tree layers of natural broad-leaved Castanopsis carlesii, its standcharacteristics such as species composition, species diversity and importance value, diameterstructure, tree height structure, volume composition, spatial structure were studyed so as toprovide reference for target tree management. The main characteristics of natural broad-leavedCastanopsis carlesii were as follow. Natural broad-leaved Castanopsis carlesii had three treelayers obviously with height more than18meters for first tree layer, height between10.1-16.9meters for second tree layer, height less than10meters for third tree layer. The dominantspecies in the research stand were Castanopsis carlesii, Schima superba, Neolitsea sericea,Elaeocarpus sylvestris, Sloanea sinensis, Castanopsis fargesii.Normal distribution and Weibulldistribution were the best model for diameter structure of each tree layer of naturalbroad-leaved Castanopsis carlesii, Gamma distribution and negative exponential distributionwere best suitable model for the diameter structure of the natural broad-leaved Castanopsiscarlesii stand.Mingling degree of stand and main tree species were medium or intensive. Thedifferiatiation value and uniform angle value of natural broad-leaved Castanopsis carlesii standwere0.48,0.75.
     (2) Target tree management incorporating selecting a small number of outstandingindividual trees as management main objective and leaving most trees in the stand under natural regulation was ecologically reasonable and ecomomic effective. The number in firsttree layer of natural broad-leaved Castanopsis carlesii was only15%of the total stand trees,but the timber volume of first tree layer had more than70%of the total stand volume. Thecharacteristics of vertical structure distribution suggested that few large diameter trees were themain part providing timber volume. Different diameter classes growth model was establishedwith “average growth plus standard deviation” method. The results showed that the standgrowth process was mainly carried out by the dominat trees which grew faster than other trees.That was, larger diameter trees whose DBH was for vigorous growth period had faster growthtrend.
     (3) Through quantitative mehods to solve the target tree density problem of naturalCastanopsis carlesii when the stand was in current stage and in harvest stage. Base on Voronoidiagram to get competition index and dominance of target trees, in the current period theaverage density of target trees of natural broad-leaved Castanopsis carlesii stand was105stems per hectare. We finally set the range of100~110stems per hectare as reasonable targettree density. In final harvest period, with use of the unite nutrition area of target tree, We gotthe vaverage signle nutrition area, average unit nutrition area and average target tree densitywere143.05m2,155.49m2,68stems per hectare respectively. By stepwise regression method,the crown wide model was established to obtain the average target tree density of Castanopsiscarlesii stand was68~74stems per hectare when target trees reached the harvest period. In theend, we got the results that the last two method for final havest period target tree density wasreasonable and in the final harvest period the appropriate target tree density of Castanopsiscarlesii stand was63~73stems per hectare.
     Based on target tree nutrition area and community completion area, we got the suitabletarget tree density was69stems per hectare.
     (4) After thinning, research resulsts showed that the growth increment and growth rate oftarget tree was significantly higher than that of non-targert tree. The fact indicated that thinningcould increase target tree growth obviously. The consequence of annual increment, averageincrement of DBH and volulme was target treeT>non-target treeNTI>non-target treeNT>non-target treeNTII>non-target treeNTIII,target treeTdiffered siginificantly with non-targerttree. The order of basal area growth rate, volume growth rate of target tree and non-target treewas non-target treeNT>target treeT>non-target treeNTI. Growth rate of basal area and volumehad the maximum2years after thinning. The growth rates of2year thinning later were notsignificant with that of3.5years thinning later, but had significantly differentce with that of1year thinning later.
     (5) By analysis basal area and volume growth rate of different management models from2008to2011, the results showed that the order of basa area growth rate of stand and each treelayer was target tree management>traditional management>control. Stand basal area growthrate of control differed significantly with that of tradional management and target treemanagement, however, the stand basal area growth rate of two latter management models hadno significant difference. The consequence of volume growth rate of stand and each tree layerwas target tree management> traditional management>control. Stand volume growth rate ofcontrol differed significantly with that of traditional management and target tree management,however, the stand volume growth rate of the two latter management models had no significantdifference.
     (6) By analysis the importance value changes before and after cutting in differentmanagement models, results showed that Castanopsis carlesii, Schima superba were still the dominantspecies. In the control test belt, the importance value of Castanopsis carlesii, Schima superba, Sycopsisdunnii, Alnus trabeculosa, Nyssa sinensis was decreasing with small tendency, importance valueof Neolitsea aurata, Sloanea changii, Symplocos lancifolia increaed. In traditional management test belt,importance value of Castanopsis carlesii, Schima superba, Neolitsea aurata, Randia cochinchinensis,Alnus trabeculosa had increased. In the target tree managemt test belt, the important value ofCastanopsis carlesii, Schima superba, Machilus chekiangensis, Alniphyllum fortune,Elacocarpus japonicas, Cryptocarya chinensis showed decreasing trendecy, the importantvalues of Neolitsea aurata,Elaeocarpus chinensis, Liquidambar formosana, Daphniphyllumoldhami, Dendropanax dentiger, Nyssa sinensis had increasing trendency.
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