用户名: 密码: 验证码:
氮磷营养及水分对冬小麦产量和品质的调控效应究
详细信息    本馆镜像全文|  推荐本文 |  |   获取CNKI官网全文
摘要
干旱是当今世界最重要的生态问题之一,也是制约农业发展的主要逆境因子,研究并揭示其对冬小麦生育中后期植株生长发育、生理特性、产量和品质特性的影响,对旱地小麦产量和品质协同提高具有重要意义。为此,结合河南西部旱地的气候特点,在自动干旱棚内遮雨条件下,设置了灌水量、施N量、施P量三个因素的水肥耦合试验,系统研究了水肥耦合对冬小麦产量形成及籽粒品质性状的调控效应,主要结论如下:
     1.在全生育期遮雨条件下,灌水量对冬小麦花后旗叶及倒二叶光合速率(Pn)、气孔导度、蒸腾速率等参数的调控效应均达显著水平(P≤0.05或P≤0.01)。灌浆前期(花后6d),旗叶Pn、气孔导度、蒸腾速率均在灌水量308.5mm达最大值,中后期(花后13d、23d)则在灌水量153.5mm时达最大值;从不同测定日期看,Pn随籽粒灌浆进程呈明显下降趋势。随施氮量增加花后6d、13d、23d旗叶Pn及气孔导度均呈先增后降趋势。从不同水肥处理组合看,以N_(105)P_(42)W_(308).5和N179.2P126W153.5最大。相关分析表明,叶片Pn与气孔导度、蒸腾速率间均呈极显著正相关(P≤0.01),而与胞间CO_2浓度则呈显著负相关(P≤0.01),说明Pn的降低主要受非气孔因素的影响。
     2.两年的研究表明,灌水量和施氮量对冬小麦籽粒蛋白质和氨基酸均有极显著调控效应(P≤0.01),存在着显著的灌水3施氮互作效应(P≤0.05或P≤0.01),而施磷对其调控效应不显著。在一定区间,籽粒蛋白质和氨基酸含量均随施氮量增加而增加,随灌水量增加而降低,但当灌水量超过282.0mm、施氮量超过179.2kg2hm~(-2)时,各指标的变化不再明显,蛋白质含量在高施氮量下略有下降。而必需氨基酸占总氨基酸含量的比例呈随灌水量增加而增加,随施氮量增加而降低的趋势,说明干旱胁迫下高氮处理不利于小麦籽粒营养品质的改善。从不同水肥处理组合看,蛋白质和氨基酸含量以处理组合N_(105)P_(42)W127最高,必需氨基酸/总氨基酸以处理组合N_(30.8)P_(126)W_(282)最高。综合从蛋白质及氨基酸产量看,以处理组合N_(179.2)P_(126)W_(282)表现最好,这与单一籽粒蛋白质及氨基酸含量高的处理组合并不一致。
     3.施氮量和施磷量对冬小麦淀粉糊化特性有显著调控效应(P≤0.05或P≤0.01);随施氮量的增加,淀粉各黏度参数呈增大趋势,施氮量为105kg2hm~(-2)时达最大值,之后增加施氮量各黏度参数又有所下降;黏度参数在施磷量为168kg2hm~(-2)时达最大值。灌水量对淀粉黏度参数调控效应不显著,但存在灌水量3施氮量的显著互作效应(P≤0.05或P≤0.01)。从不同水肥处理组合看,淀粉糊化特性以处理组合N_(105)P_(168)W_(217.5)最优。相关分析表明,淀粉黏度参数与籽粒产量在2009—2010年度呈显著正相关,而在2010—2011年度相关不显著,说明产量与淀粉品质间存在着协同提高的可能性。
     4.灌水量对冬小麦产量及其构成因素的调控效应显著(P≤0.01)。随灌水量增加,总茎数及最终成穗数增加,在灌水量282.0~308.5mm时达最大値;穗粒数及产量在灌水量308.5mm达最大值,千粒重在灌水量282.0mm最大。在施氮量30.8~179.2kg2hm~(-2)区间,总茎数随施氮量增加呈上升趋势;产量在0.0~105.0kg2hm~(-2)间呈增加趋势。从水肥耦合的效应看,组合N_(179.2)P_(126)W_(282)对总茎数的调控效应较大,产量、穗粒数在组合N_(105)P_(42)W_(308).5时最大,千粒重以N_(30.8)P_(126)W_(282)达最大值。
     5.在全生育期遮雨条件下,灌水量对冬小麦产量及水分利用率的调控效应显著(P≤0.01),冬小麦水分利用效率在灌水量308.5mm达最大值,在灌水量127.0mm时为最小值,即随灌水量增加呈增加趋势,表明在全生育期遮雨条件下,因小麦生育后期的干旱程度不同,导致灌水量增加的情况下小麦的籽粒产量仍然大幅提高,灌水量仍然是水分利用效率提高的限制因素;随施氮量和施磷量增加水分利用效率呈先增后降再增的趋势,分别在施氮量210.0kg2hm~(-2)、施磷量84.0kg2hm~(-2)达最高值,在施氮量0.0kg2hm~(-2)及施磷量0.0kg2hm~(-2)最低。且籽粒产量及水分利用效率均在水肥处理组合N_(105)P_(42)W_(308).5最高值。籽粒产量与水分利用效率间呈极显著正相关(P≤0.01),说明在一定区间,高水分利用率处理小麦产量亦较高。
Nowadays drought is one of the most important ecological problems in the world,and the most serious stress factors restricting development of agriculture. A betterunderstanding of drought and fertilizer effects on growth and development,physiological characters, grain yield and quality traits of winter wheat is veryimportant to improve both grain yield and quality. To evaluate the effects ofwater-fertilizer coupling on yield formation and grain quality traits of winter wheat,experiment was carried out in rain-sheltered condition with factors of water, nitrogenand phosphorus fertilization in a typical arid climate in middle China for two years.The results were as follows.
     1. Irrigation had significant effects on the photosynthetic rate, stomatalconductance, transpiration rate in flag leaf and top second leaf in rain-shelteredcondition during the whole growth of wheat (P≤0.05or≤0.01). Photosynthetic ratereached the highest value at irrigation amount of308.5mm in the day of6days afteranthesis (DAA), and at the amount of153.5mm both in13and23DAA. While thephotosynthetic rate in flag leaf decreased sharply during grain filling period. With theincrease of nitrogen application, the photosynthetic rate and stomatal conductance inflag leaf increased and then decreased on6,13,23DAA. N_(105)P_(42)W_(308).5andN179.2P126W153.5obtained the highest values of photosynthetic traits among allcombinations of treatment. Correlation analysis indicated that, the photosynthetic ratein leaf significant positively correlated with stomatal conductance and transpirationrate (P≤0.01), while significant negatively correlated with intercellular CO_2concentration (P≤0.01), implying that decline of photosynthetic rate was mainlycaused by non-stomatal factors.
     2. The results of two year study indicated that irrigation and nitrogen applicationhad significant effects on grain protein and amino acids contents (P≤0.01) of winterwheat. The interaction of irrigation and nitrogen fertilization also presented significanteffects (P≤0.05or≤0.01). No significant effect of P fertilization on protein and aminoacids contents was observed. Protein and amino acid contents in wheat grainsincreased with increasing nitrogen fertilization rate, but dropped with increasing ofirrigation amount. However, as irrigation amount was above282.0mm, and nitrogenfertilization rate was above179.2kg·hm~(-2), the changes became not less obvious.Protein content decreased as the nitrogen fertilization rate above179.2kg·hm~(-2). The trend of ETT/ATT was contrary to that of amino acid content. It indicated that highnitrogen treatment with drought stress went against of the improvement of grainquality. The contents of protein and amino acids in wheat grains reached the highest intreatment of N_(105)P_(42)W127with nitrogen fertilization rate of105kg(N)·hm~(-2),phosphorus fertilization rate of42kg(P2O5)·hm~(-2)and irrigation amount duringgrowing season of127mm. While ETT/ATT reached the highest in treatment ofN_(30.8)P_(126)W_(282)with nitrogen fertilization rate of30.8kg(N)·hm~(-2), phosphorusfertilization rate of126kg(P2O5)·hm~(-2)and irrigation amount during growing seasonof282mm. Overall, N_(179.2)P_(126)W_(282)with nitrogen fertilization rate of179.2kg(N)·hm~(-2), phosphorus fertilization rate of126kg(P_2O_5)·hm~(-2)and irrigation amountduring growing season of282mm was the best treatment.
     3. Application of nitrogen and phosphorus had significant effects on starchpasting properties (P≤0.05or P≤0.01) of winter wheat. The main starch viscosityparameters (peak viscosity, through viscosity and final viscosity) were increasedsignificantly when nitrogen application increased from0kg·hm-2to105kg·hm-2. Andthe starch viscosity parameters reached the highest values at the phosphatic amount of168kg·hm-2. No significant effect of irrigation on starch pasting properties wasobserved. But interaction of irrigation×nitrogen had significant effects on someparameters of starch pasting property (P≤0.05or≤0.01). Overall, the combination oftreatments N_(105)P_(168)W_(217.5), with nitrogen rate of105kg(N)·hm~(-2), phosphorus rate of168kg(P2O5)·hm~(-2)and irrigation amount of217.5mm, obtained the best starchquality in the experiment. Correlation analysis indicated that, starch viscosityparameters had positive significant correlation with grain yield in crop season of2009~2010,but had not correlation in crop season of2010~2011. And it indicatedthat possibility of improving both grain yield and starch content was exist.
     4. Irrigation had significant effects on grain yield and its components (P≤0.01) ofwinter wheat. Both the culm population and ear number increased with enhance ofirrigation amount, and reached the highest value at irrigation amount of282.0~308.5mm. Grain number per ear and grain yield reached the highest values atirrigation amount of308.5mm, while thousand kernel weight (TKW) at irrigationamount of282.0mm. The culm population increased with nitrogen applicationimprovement from30.8kg·hm-2to179.2kg·hm-2. Grain yield increased when nitrogen application increased from0kg·hm-2to105kg·hm-2. The treatment of N_(179.2)P_(126)W_(282)obtained the largest culm population, for grain number per ear and grain yield,treatment of N_(105)P_(42)W_(308).5,and for TKW, N_(30.8)P_(126)W_(282)obtained the highest values,respectively.
     5. Irrigation had significant effect on grain yield and water utilization efficiencyin rain-sheltered condition during the whole growth of wheat (P≤0.01). The waterutilization efficiency reached the highest value at irrigation amount of308.5mm,lowest at127.0mm. It indicated that irrigation amount was still a limiting factor forincreasing the water utilization efficiency in arid area. The water utilization efficiencyreached the highest at nitrogen rate of210.0kg·hm~(-2), phosphorus rate of84.0kg·hm~(-2),while got the lowest value when no nitrogen and phosphorus application. Thecombination of treatments N_(105)P_(42)W_(308).5obtained the highest value for both grainyield and water utilization efficiency. Grain yield had significant positive correlationwith water utilization efficiency (P≤0.01). So the treatment of higher water utilizationefficiency always had higher grain yield.
引文
[1]王建进,高秀兰,李守霞,等.旱地小麦高产途径与栽培技术.安徽农学通报,2008,14(2):107-108.
    [2]山仑,张岁歧.能否实现大量节约灌溉用水?我国节水农业现状与展望.资源环境与发展,2006,(1):1-4.
    [3]康绍忠,张富仓,梁银丽,等.土壤水分和CO2浓度增加对小麦、玉米、棉花蒸散、光合及生长的影响.作物学报,1999,25(1):55-63.
    [4] Wang T, Zhang X, Li C. Growth, abscisic acid content, and carbon isotope composition in wheat cultivarsgrown under different soil moisture[J]. Biologia Plantarum,2007,51(1):181-184.
    [5] Kage H, Kochler M, Stützel H. Root growth and drymatter partitioning of cauliflower under drought stressconditions: measurement and simulation. Eur J Agron,2004,20:379-394.
    [6] Kozlowski T T, Pallardy S G. Acclimati on and adaptive responsesf woody plants to environmentalstresses. Bot Rev,2002,68:270-334.
    [7] Ober E S, Setter T L, Madison J T, et al. Influence of water deficiton maize endospermdevelopment.Enzyme activities and RNA transcripts of starch and zein synthesis, abscisic acid, and celldivision. Plant Physiology,1991,97:154-164.
    [8] Saini H S, Westgate M E. Reproductive development in grain cropsduring drought. Advances in Agronomy,2000,68:59-95.
    [9] Brocklehurst P A. Factors controlling grain weight in wheat.Nature,1977,266:348-349.
    [10] Nicolas M E, Lambers H, Simpson R J, et al. Effect of drought on metabolism and partitioning of carbon intwo wheat varieties differing in drought to lerance. Annals of Botany,1985,55:727-747.
    [11] Keck R W, Boyer J S, Chloroplast response to low leaf water potentials.Ⅲ Differing inhibition of electrontransport and photophosphorylation. Plant Physiol,1974,53:474-479.
    [12] Younis H M, Boyer J S, Covindjee. Conformation and activity of chloroplast coupling factor exposed tolow chemical potential of water in cells. Biochem Biophys Acta,1979,548:328-340.
    [13] Kaiser W M,.Effects of water deficit on photosynthetic capacity. Physiol Plant,1987,71:142-149.
    [14]王志敏.小麦籽粒蛋白质贮积的生理学研究进展.国外农学-麦类作物,1996(4):23-26.
    [15]郭天财,姚战军,王晨阳,等.水肥运筹对小麦旗叶光合特性及产量的影响.西北植物学报,2004,24(10):1786-1791.
    [16]刘建福,汤青林,倪书邦,等.水分胁迫对澳洲坚果叶绿素a荧光参数的影响.华侨大学学报,2003,7(3):305-309.
    [17]许振柱,李长荣.土壤干旱对冬小麦生理特性和干物质积累的影响.干旱地区农业研究,2000,18(1):113-123.
    [18]梁新华,许兴,徐兆桢,等.干旱对春小麦叶绿素a荧光动力学特征及产量间关系的影响.干旱地区农业研究,2001,19(3):72-77.
    [19]王小燕.施氮量和土壤水分对小麦碳氮代谢和产量与品质形成的影响.山东农业大学,2005.
    [20]张其德,刘合芹,张建华,等.限水灌溉对冬小麦旗叶某些光合特性的影响.作物学报,2000,26(6):869-873.
    [21]张忠学,于贵瑞.不同灌水处理对冬小麦生长及水分利用效率的影响.灌溉与排水学报,2003,22(2):36-39.
    [22]吴乃元,梁丰香,张衍华,等.有限水分胁迫对小麦生长状况的影响及合理灌溉的土壤相对湿度指标.应用气象学报,2000,11(11):170-177.
    [23]王晨阳,郭天财,彭羽,等.花后灌水对小麦籽粒品质性状及产量的影响.作物学报,2004,30(10):1031-1035.
    [24]吕金印,山仑.水分亏缺对小麦碳同化物的动员与分配.核农学报,2002,16(4):228-231.
    [25]张永丽,于振文.灌水量对不同小麦品种籽粒品质、产量及土壤硝态氮含量的影响田.水土保持学报.2007,21(5):155-158.
    [26]王立秋,靳占忠,曹敬山,等.水肥措施对冬小麦产量及其构成因素的影响.麦类作物学报,1996,6:40-43.
    [27]许振柱,于振文.限量灌溉对冬小麦水分利用的影响.旱地区农业研究,2003,21(1):6-10.
    [28]王俊儒,李生秀.不同生育时期水分有限亏缺对冬小麦产量及其构成因素的影响.西北植物学报,2000,20(2):193-200.
    [29]赵世伟,管秀娟,吴金水.不同生育期干旱对冬小麦产量及水分利用率的影响.灌溉排水,2001,20(4):56-59.
    [30]梅雪英,严平,王风文,等.水分胁迫对冬小麦根系生长及产量的影响叨.安徽农业科学,2003,31(6):962-964.
    [31]兰霞,周殿玺,兰林旺.灌溉制度对冬小麦产量结构形成与产量物质来源的影响.中国农业大学学报,2001,6(1):17-22.
    [32]於俐,于强.水分胁迫对冬小麦物质分配及产量构成的影响.地理科学进展,20041(1):105-111.
    [33]胡继超,曹卫星,姜东.小麦水分胁迫影响因子的定量研究I干旱和渍水胁迫对光合、蒸腾及干物质积累与分配的影响.作物学报,2004,4(4):315-320.
    [34]刘庚山,郭安红.人工控制有限供水对冬小麦根系生长及土壤水分利用的影响.中国农业大学学报,2001,6(1):17-22.
    [35]付雪丽,王晨阳,郭天财,等.水氮互作对小麦籽粒蛋白质、淀粉含量及其组分的影响[J].应用生态学报,2008,19(2):317–322
    [36]李卫华,张东海.小麦籽粒形成期间氨基酸含量的平衡性分析[J].种子,2000(2):21–23
    [37]安成立,张改生,高翔,等.不同生态环境对强筋小麦品质的影响[J].中国生态农业学报,2006,14(1):3436
    [38]章练红,王绍中,李运景,等.小麦品质生态研究概述与展望.国外农学—麦类作物,1994,6:42-44.
    [39] Pheloun Pc,Siddique KHM. Contribution of stem dry matter to grain yield in wheat cultivars,Aust J PlantPhysiol.1991,18:53-64.
    [40]王月福,陈建华,曲健磊,等.土壤水分对小麦籽粒品质和产量的影响.莱阳农学院学报,2002,19(1):7-9.
    [41]许振柱,于振文,王东,等.灌溉条件对小麦籽粒蛋白质组分积累及其品质的影响.作物学报,2003,29(5):682-687.
    [42]赵广才,周阳,常旭虹,等.氮磷钾硫对冬小麦产量及加工品质的调节效应.植物遗传资源学报,2005,6(4):423-426.
    [43]刘玲,沙奕卓,白月明.中国主要农业气象灾害区域分布与减灾对策.自然灾害学报.2003,12(2):92-97.
    [44]范雪梅,姜东,戴廷波,等.花后干旱和渍水逆境下氮素对小麦籽粒产量和品质的影响.植物生态学报,2006,30(1):71-77.
    [45]毛凤梧,赵会杰,段藏禄.潮土麦田施磷对小麦品质的影响初探.河南农业大学学报,2001,35(4):400-402.
    [46]杨胜利,冯荣成,马利民.后期灌溉对强筋小麦产量和品质的影响.河南职业技术师范学院学报,2002,30(3):4-6.
    [47]胡祥辉,胡茂兴.小麦栽培生理[M].上海科学技术出版社,1984,269-259
    [48]翟丙年,孙春梅,王俊儒等.氮素亏缺对冬小麦根系生长发育的影响.作物学报,2003,29(6):913~918
    [49]任书杰,张雷明,张岁岐等.氮素营养对小麦根冠协调生长的调控.西北植物学报,2003,23(3):395~400
    [50]张殿顺,董刘树,庆翔云等.不同施氮水平对春小麦生长发育及其氮素代谢指标的影响.华北农学报,2006,21(2)
    [51]张丽霞.不同氮肥水平对冬小麦光合产物分配格局及生理生态特性的影响.2008,硕士论文
    [52]张会绒.不同施氮水平对小麦幼苗性状的影响.陇东学院学报,2010,05
    [53] Campbell CA, Cameron DR, Nicholaichuk W, et al. Effects of fertilizer N and soil moisture on growth, Ncontent and moisture use by spring whent. Canadian Journal of Soil Science,1977,57:289-310
    [54] Villalobos F J,Fereres E. Evaporation measurements beneath corn, cotton,and sunflower canopies.Agronomy Journal,1990,82:1153-1159
    [55]孟繁静,刘宏道,苏业瑜.植物生理生化[M].北京:中国农业出版社1995,36-37
    [56]张岁岐,李秧秧.施肥促进作物水分利用机理及对产量影响的研究.水土保持研究,1996,3(1),185-191
    [57]赵波海,林琪,刘义国,等.氮磷肥配施对超高产冬小麦灌浆期光合日变化及产量的影响.应用生态学报,2010,21(10),2545-2550
    [58]尹光华等.水肥耦合条件下春小麦叶片的光合作用.兰州大学学报,2006,42(1)
    [59]孙旭生,林琪,李玲燕,等.氮素对超高产小麦生育后期光合特性及产量的影响.植物营养与肥料学报,2008,05
    [60]林同保,崔国金,王志强,等.拔节期水氮运筹对冬小麦籽粒灌浆期叶片光合特性及水分.麦类作物学报,2009,29(4),627-631-
    [61]潘庆民,于振文,王月福,等.追氮时期对小麦旗叶中蔗糖合成与籽粒中蔗糖降解的影响.中国农业科学.2002,35(7):771-776
    [62]王月福,于振文,李尚霞.氮素营养水平对冬小麦氮代谢关键酶活性变化和籽粒蛋白质含量的影响.作物学报,2002,(6):743-748.
    [63]赵广才,张保明,王崇义.不同类型冬小麦氮素积累及施氮对策探讨.作物学报,1998,24(6):894-898.
    [64] Dhugga KS,Waines J G.Analysis of nitrogen accumulation and use in bread and durum wheat.CropScience,1989,29:1232-1239
    [65]王月福,于振文,李尚霞.土壤肥力和施氮量对小麦氮素吸收运转及籽粒产量和蛋白质含量的影响.生态学报,2003,14(11):1868-1872.
    [66]王月福,于振文,李尚霞.氮素营养水平对小麦开花后碳素同化、运转和产量的影响.麦类作物学报,2002,22(2):55-59.
    [67]朱新开,郭文善,周正权.氮肥对中筋小麦扬麦10号氮素吸收、产量和品质的调节效应.中国农业科学,2004,37(12):1831-1837.
    [68]赵俊晔,于振文.高产条件下施氮量对冬小麦氮素吸收分配利用的影响.作物学报,2006a,32(4):484-490.
    [69]张洪程,许柯,戴其根.超高产小麦吸氮特性与氮肥运筹的初步研究.作物学报,1998,24(6):935-940.
    [70]石玉,于振文,王东,等.施氮量和底追比例对小麦氮素吸收转运及产量的影响.作物学报,2006,32(12):1860-1866.
    [71]李秀章,陈祥龙,徐立华,等.氮素营养水平对小麦后移栽棉氮代谢的影响.棉花学报,1994,6(4):223-228.
    [72]关义新,林葆,凌碧莹.光、氮及其互作对玉米幼苗叶片光合和碳、氮代谢的影响.作物学报,2000,26(6):806-812.
    [73]王小纯,熊淑萍,马新明,等.不同形态氮素对专用型小麦花后氮代谢关键酶活性及籽粒蛋白质含量的影响.生态学报,2005.4:802-806
    [74]王月福,于振文,李尚霞,等.土壤肥力和施氮量对小麦根系氮同化及子粒蛋白质含量的影响.植物营养与肥料学报,2003,9(1):39-44
    [75]隋益虎,朱世东,张子学,等.环境因子对苋菜碳、氮代谢关键酶及抗氧化酶活性的影响.生态学杂志,2005,24(8):925-929
    [76]巨晓棠,张福锁.关于氮肥利用率的思考[J].生态环境,2003,12(2),192-197
    [77]赵会杰,薛延丰,董中东,等.密度及追氮时期对大穗型小麦旗叶及子粒碳水化合物代谢的影响.河南农业大学学报,2002,38(1):1-4
    [78]赵俊晔,于振文.施氮量对小麦旗叶光合速率和光化学效率、籽粒产量与蛋白质含量的影响.麦类作物学报,2006b,26(5):92-96.
    [79]宁东峰,李志杰,孙文彦,等.节水灌溉对黄淮海地区冬小麦水分消耗与光合特性的影响.植物营养与肥料学报,2010,16(4)
    [80]彭永欣.小麦栽培与生理.南京:东南大学出版社,1992
    [81]刘安勋.氮肥对小麦籽粒氨基酸的贡献.青海农林科技,2001,3(1)
    [82]杜朝,杨学举,刘桂茹等.小麦面粉淀粉特性与烘烤品质关系的研究.河北农业大学学报,2002,25(4),29-33
    [83]王晨阳,马冬云,郭天财等.不同水_氮处理对小麦淀粉组成及特性的影响.作物学报,2004,30(8):739-744.
    [84]黄明丽,邓西平,白登忠. N、P营养对旱地小麦生理过程和产量形成的补偿效应研究进展.麦类作物学报,2002,22(4):74-78.
    [85] Steven J,Crafts-Brander ME,Salvucci JL,etal. Phosphorus nutrion influence on plant growthand nonstructural carbohydrate accumulation in Tobacco. Crop Sci.,1990,30:509-604.
    [86] Jacob J,Lawlor DW. Stormatal and mesophyll limitation of lhotosynthesis in phosphatedeficient sunflower,maize,wheat plant. J.Exp.Bot.,1991,42:1003-1011.
    [87]张振清,夏叔芳.无机磷对叶片淀粉和蔗糖积累的影响。植物生理学报,1982,8(4):385-391.
    [88] Rao IM,Arthur L. Fredeen,Norman Terry. Leaf phosphate status,photosynthesis,and carbonpartitioning in sugar beet. Plant Physiol.,1990,92:29-36.
    [89] Hideaki U,Kousuke S. Phosphate deficiency in Maize enzyme activities. Plant Cell Physiol,1991,32:1313-1317.
    [90]金善宝.中国小麦.北京:中国农业出版社,1996.
    [91]王旭东,于振文,石玉,等.磷对小麦旗叶氮代谢有关酶活性和籽粒蛋白质含量的影响.作物学报,2006,32(3):339-344.
    [92]毛凤梧,赵会杰,徐立新等.水肥运筹对小麦品质形成的调控效应阴.河南农业大学学报,2001,35(1):13-15.
    [93]吴克宁,赵彦锋,吕巧灵,等.潮土区灌浆水和施磷对冬小麦光合作用和产量的影响.植物营养与肥料学报,2002,8(4):428-434.
    [94]张起刚,王声斌,彭根元,等.用15N示综研究灌溉水平和氦磷配比对沙地小麦生长和氮素平衡的影响.中国农业大学学报,1996,1(3):33-38.
    [95]岳寿松,于振文.磷对冬小麦后期生长及产量的影响.山东农业科学,1994,(1):13-15.
    [96] Sweeny D W,Granade GV. Evereyer MG,Whitney DA. Phosphorus,potassium,chloride andfungicide effects on wheat yield and leaf rust.Journal of Plant Nutrition,2000,23(9):1267-1281.
    [97]彭正萍,李春俭,门明新.缺磷对不同穗型小麦光合生理特性和产量的影响.作物学报,2004,30(8):739-744
    [98]王兰珍,米国华,陈范骏,张福锁.不同产量结构小麦品种对缺磷反应的分析.作物学报,2003,29(6):867-870.
    [99]王兰珍,米国华,陈范骏,等.冬小麦品种的磷营养效率鉴定.中国农业大学学报,2003,8(5):69-73.
    [100]张礼军,张恩和,黄高宝.灌溉与供磷对小麦-玉米-马铃薯间作系统小麦产量和品质的调控效应.麦类作物学报,2007,27(4):687-692.
    [101]王旭刚,郝明德,李建民,等.氮磷配施对旱地小麦产量和吸肥特性的影响.西北农林科技大学学报:自然科学版,2007,35(2):138-142.
    [102]张国平,王中琪.几种农艺因素影响小麦籽粒蛋白质含量的数学模型研究.生物数学学报,1995,10(4):148-152.
    [103]王旭东,于振文.施磷对小麦产量和品质的影响.山东农业科学,2003(6):35-36.
    [104]季书勤,郭瑞,赵淑章,等.氮磷钾、灌水、喷洒杀菌剂对强筋小麦产量和品质的影响.华北农学报,2006,21(4):82-86.
    [105] Batten GD. A review of phosphorus efficiency in wheat.Plant and Soil,1992,146:163-168.
    [106]曹卫星,郭文善,王龙俊,等.小麦品质生理生态及调优技术.北京:中国农业出版社,2005.
    [107]赵广才,舸中虎.刘利华,等.肥水调控对强筋小麦中优9507品质与产量协同提高的研究.中国农业科学,2004,3:351-356.
    [108]姜宗庆,封超年,黄联联,等.施磷量对小麦物质生产及吸磷特性的影响.植物营养与肥料学报,2006,12(5):628-633.
    [109]孙慧敏,于振文,颜红,等.不同土壤肥力条件下施磷量对小麦产量、品质和磷肥利用率的影响.山东农业科学,2006(3):45-47.
    [110]张睿,刘党校.氮磷与有机肥配施对小麦光合作用及产量和品质的影响.植物营养与肥料学报,2007,13(4):543-547.
    [111]姜东,戴廷波,荆奇,等.氮磷钾肥长期配合施用对冬小麦籽粒品质的影响.中国农业科学,2004,37(4):566-571.
    [112]赵淑章,王绍中,付国占,等.氮、磷、钾对强(弱)筋小麦品质的效应研究.中国农学通报,2004,20(1):152-154.
    [113]吕殿青、张文孝、谷洁等.渭北东部旱塬氮磷水三因素交互作用研究[M].北京:中国农业科技出版社,1995:286-292
    [114]尹光华,刘作新,李桂芳等.辽西半干旱区春小麦氮磷水耦合产量效应研究.农业工程学报,2005,21(1),41-45
    [115]王立秋,曹敬山,靳占忠等.春小麦产量及其品质的水肥效应研究.干旱地区农业研究.1997,15(1)
    [116]王晨阳,马冬云,郭天财,等.不同水、氮处理对小麦淀粉组成及特性的影响[J].作物学报,2004,30(8):739744
    [117]张依章,张秋英,孙菲菲,等.水肥空间耦合对冬小麦光合特性的影响.干旱地区农业研究,2006,24(2)
    [118]董树连,王卫国,李春茂,等.旱地高产小麦光合速率与产量变化的研究.莱阳农学院学报,2000,17(3):194-195.
    [119]黄令峰,林琪,刘义国,等.限量补灌对旱地高产田小麦光合和产量的影响.中国农学通报,2008,24(2),470-474
    [120]郭天财,方保停,王晨阳,等.水分调控对小麦旗叶叶绿素荧光动力学参数及其产量的影响.干旱地区农业研究,2005,23(2),6-10
    [121]赵春江,郭晓维,张其德,等.不同穗型冬小麦叶片荧光诱变及光谱特性.中国农业科学,1999,32(2),43-46
    [122]范雯雯,王志强,林同保,等.不同土壤肥力下旱后复水对冬小麦光合特性及水分利用效率的影响,麦类作物学报,2010,2
    [123]李玉欣,师长海,乔匀周,等.不同水分条件下作物蒸腾效率的比较研究.干旱地区农业研究,2010,6,
    [124]戚龙海,党廷辉,陈璐.黄土旱塬冬小麦水分利用效率及相关生理特性研究.中国农学通报,2009,6
    [125]鞠正春,于振文.追施氮肥时期对冬小麦旗叶叶绿素荧光特性的影响.应用生态学报,2006,3
    [126]王佩玲,许育彬,宋淑英,等.大气CO_2浓度倍增和施氮对冬小麦光合及叶绿素荧光特性的影响.西北植物学报,2011,1
    [127]姜东燕,于振文.不同灌溉量对冬小麦旗叶叶绿素荧光特性的影响.安徽农业科学
    [128] Anjum F M, Ahmad I, Sheikh M A, et al. Amino acid composition of spring wheat and lossesof lysine during chapatti baking[J]. Journal of Food Composition and Analysis,2005,18:523–532
    [129]曹广才,王绍中,小麦品质生态,北京:中国科学技术出版社,1994:89-92.
    [130]张林生.小麦种子氨基酸的评价[J].国外农学—麦类作物,1996,11(3):28–30
    [131] Singh A K, Jain G L. Effect of sowing time, irrigation and nitrogen on grain yield and qualityof durum wheat (Triticum dunm)[J]. Indian Journal of Agricultural Sciences,2000,70(8):532–533
    [132]田志会,孙彦,郭玉琴.主要生态因素对小麦营养品质及烘烤品质的影响.北京农学院学报,2000,15(2):67-71.
    [133]王立秋.小麦面食蒸煮品质研究动态,国外农学—麦类作物,1994,6:45-47.
    [134]汪永钦,杨海鹰,刘荣花,等.试论冬小麦籽粒品质与气象条件的关系.中国农业气象,1990,11(2): l-7.
    [135]王晓燕,卢少源,荣广哲,等.第三次全国面包小麦品种(系)品质分析,河北农业大学学报,1997,20(2):1-5.
    [136]白莉萍,林而达,饶敏杰.不同试点灌溉方式对冬小麦产量和品质性状的影响[J].生态学报,2005,25(4):917–922
    [137]赵广才,常旭红,陈新民,等.不同施肥灌水处理对不同小麦品种产量和品质的影响[J].植物遗传资源学报,2007,8(4):447–450
    [138]王月福,姜东,于振文,等.施氮水平对小麦籽粒产量和蛋白质含量的影响及其生理基础[J].中国农业科学,2003,36(5):513–520
    [139]陆欣春,田霄鸿,杨习文,等.氮锌配施对不同冬小麦品种产量及锌营养的影响[J].中国生态农业学报,2010,18(5):923–928
    [140]翟丙年,李生秀.水氮配合对冬小麦产量和品质的影响[J].植物营养与肥料学报,2003,9(1):26–32
    [141]曹广才.华北小麦品质[M].北京:中国农业出版社,2001:207–221
    [142]王晨阳,马冬云,朱云集,等.小麦不同水氮运筹对面条煮制品质的影响[J].中国农业科学,2004,34(2):256–262
    [143]李焕春,高炳德,妥德宝,等.氮肥品种及施用时期对春小麦籽粒氨基酸含量的影响[J].中国土壤与肥料,2009(1):12–15
    [144]张林生,张保军,汪沛洪.等.施氮水平对小麦籽粒发育过程中氨基酸含量的影响[J].西北植物学报,2002,22(3):646–650
    [145]姜东燕,于振文.灌溉量对冬小麦产量和土壤硝态氮含量的影响[J].土壤通报,2008,39(3):703–705
    [146]王绍中,李春喜,罗艳蕊,等.基因型和地域分布对小麦籽粒氨基酸含量影响的研究[J].西北植物学报,2001,21(3):437–445
    [147]孔令聪,汪建来,曹承富,等.主要栽培措施对中筋小麦皖麦44产量和品质的影响[J].麦类作物学报,2004,24(4):84–87
    [148]池忠志,赵广才,常旭虹,等.施氮量对冬小麦籽粒蛋白质及其组分含量和比例的影响[J].麦类作物学报,2006,26(6):65–69
    [149]姚大年,李保云,梁荣奇,等.基因型和环境对小麦品种淀粉性状及面条品质的影响[J].中国农业大学学报,2000,5(1):6368
    [150]马冬云,郭天财,王晨阳,等.施氮水平对小麦籽粒淀粉粒度分布及淀粉粒糊化特性的影响[J].西北农业学报,2010,19(11):4347
    [151]宋健民,刘爱峰,李豪圣,等.小麦籽粒淀粉理化特性与面条品质关系研究[J].中国农业科学,2008,41(4):272279
    [152]刘建军,赵振东,徐亚洲,等.淀粉质量与面条煮面品质的关系[J].山东农业科学,1999(6):4851
    [153] Toyokawa H, Rubenthaler G L, Powers J R, et al. Japanese noodle qualities. Ⅱ. Starchcomponents[J]. Cereal Chemistry,1989,66(4):387391
    [154] Crosbie G B, Miskelly D M, Dewan T. Wheat quality for Japanese flour milling and noodleindustries[J]. W A Journal of Agriculture,1990,31:8394
    [155]方保停,何盛莲,郭天财,等.水分调控对两种筋力型小麦品种籽粒淀粉糊化特性的影响[J].水土保持学报,2005,19(6):162165
    [156]蔡瑞国,尹燕枰,张敏,等.氮素水平对藁城8901和山农1391籽粒品质的调控效应[J].作物学报,2007,33(2):304310
    [157]王晨阳,何英,方保停,等.小麦籽粒淀粉合成、淀粉特性及其调控研究进展[J].麦类作物学报,2005,25(1):109114
    [158]阎俊,张勇,何中虎.小麦品种糊化特性研究[J].中国农业科学,2001,34(1):913
    [159]张学林,郭天财,朱云集,等.河南省不同纬度生态环境对三种筋型小麦淀粉糊化特性的影响[J].生态学报,2004,24(9):20502055
    [160]王晨阳,郭天财,马冬云,等.环境、基因型及其互作对小麦主要品质性状的影响[J].植物生态学报,2008,
    [161]马冬云,朱云集,郭天财,等.基因型和环境及其互作对河南省小麦品质的影响及品质稳定性分析[J].麦类作物学报,2002,22(4):1318
    [162]冯伟,李晓,郭天财,等.水氮运筹对两种穗型冬小麦品种淀粉糊化特性的影响[J].水土保持学报,2005,19(5):186190
    [163]张美微,王晨阳,贺德先,等.环境和氮磷肥对强筋小麦品种郑麦9023淀粉糊化特性的影响[J].麦类作物学报,2010,30(5):905909
    [164]王晨阳,冀天会,郭天财,等.干旱胁迫对春小麦淀粉糊化特性的影响[J].河南农业科学,2008(8):3237
    [165]熊瑛,李友军.氮、磷、钾对不同筋型小麦产量和品质的影响[J].河南科技大学学报:自然科学版,2005,26(3):5861
    [166] Reddy I, Seib P A. Paste properties of modified starches from partial waxy wheats[J]. CerealChemistry,1999,76(3):341349Rodriguez D,Andrade FH,Goudriaan J.Effects of phosphorusnutrition on tiller emergence in wheat.Plant and Soil,1999,209:283-295.
    [167]李友军,付国占,刘丰明,等.拔节期重施氮肥对小麦群体质量和产量的影响.麦类作物学报,1997,17(5),41-45
    [168]叶优良,王桂良,朱云集,等.施氮对高产小麦群体动态_产量和土壤氮素变化的影响.应用生态学报,2010,21(2)351-358
    [169]刘强,葛鑫,于松溪,等.氮肥运筹对强筋小麦济南17群体结构和产量的影响.耕作与栽培,2003年05期
    [170]许昌惠,徐珍秀,余毓君,等.小麦群体发育特性的研究.华中农业大学学报,1982,3,1-21
    [171]周瑞.不同水肥耦合处理下冬小麦光合产物分配格局和动态研究.2009,硕士论文
    [172]杨静敬,蔡焕杰,王健,等.灌水处理对冬小麦生理生长特性等的影响研究.灌溉排水學報,2009,28期(1)
    [173]张岁岐,山仑.作物水分利用效率及其研究进展.干旱地区农业研究,2002,20(4),1-5
    [174]张洋,翟丙年.水分胁迫条件下氮素营养对不同冬小麦基因型的生理效应.麦类作物学报,2010,30(3),492-495
    [175]郑成岩,于振文,张永丽,等.不同施氮水平下灌水量对小麦水分利用特征和产量的影响.应用生态学报,2010,21(11),2799-2805
    [176]赵广才,刘利华,张艳,等.肥料运筹对超高产小麦群体质量_根系分布_产量和品质的效应.华北农学报,2002,17(4),82-87
    [177]赵雪飞.水氮运筹对冬小麦群体物质生产特性和产量形成的影响.硕士论文,2009
    [178]宁东峰,李志杰,孙文彦,等.限水灌溉下施氮量对冬小麦产量_氮素利用及氮平衡的影响.植物营养与肥料学报,2010,16(6),1312-1318
    [179]姚金保,杨学明,马鸿翔,等.拔节期追施氮肥对宁麦16产量和品质的影响.麦类作物学报,2010,30(4),727-730
    [180]段敏,同延安,魏样.不同施肥条件下冬小麦氮素吸收_转运及累积的研究.麦类作物学报,2010,30(3),464-468
    [181]王小燕,沈永龙,高春宝,等.氮肥后移对江汉平原小麦籽粒产量及氮肥偏生产力的影响.麦类作物学报,2010,30(5)
    [182]尹光华,刘作新,李桂芳,等.水肥耦合对春小麦水分利用效率的影响.水土保持学报,2004,18(6),156-158
    [183]张洁,姚宇卿,吕军杰,等.水肥耦合对小麦产量及淀粉特性的影响.中国粮油学报,2011,26(12),1-4
    [184]张正斌,徐萍,董宝娣,等.水分利用效率——未来农业研究的关键问题.世界科技研究与发展,2005年01期
    [185]崔欢虎,王娟玲,马步州,等.茬口和灌水对小麦产量及水分利用效率的影响[J].中国生态农业学报,2009,17(3):479–483
    [186]谭念童,林琪,姜雯,等.限量灌溉对旱地小麦旗叶光合特性日变化和产量的影响.中国生态农业学报,2011,19(4):805–811
    [187]董宝娣,刘孟雨,张正斌.不同灌水对冬小麦农艺性状与水分利用效率的影响研究.中国生态农业学报,2004,12(1):140–143
    [188]武天云,王生录,邓娟珍,等.陇东旱塬地区冬小麦水肥效应耦合模拟研究.:西北农业学报,1995,4(2),69-72
    [189]汪玉磊,杨劲松,杨晓英.水肥耦合对冬小麦产量、品质和氮素利用的影响研究.灌溉排水学报,2008,27(6),31-33
    [190]周立峰,冯浩.新型有机无机复合肥对冬小麦水肥利用效率的影响.节水灌溉,2011年10期
    [191]孟晓瑜,底墒和氮磷用量对旱地冬小麦产量形成、养分水分利用的影响.2011,硕士论文
    [192] Dobermann A. Nitrogen use efficiency-state of the art Paper of the IFA international Workshopon Enhanced-Efficiency Fertilizers Fankfurt,Germany.2005
    [193]马兴华,于振文,梁晓芳,等.施氮量和底追比例对小麦氮素吸收利用及子粒产量和蛋白质含量的影响.植物营养与肥料学报,2006,12(2),150-155
    [194]苏佩,蒋纪云,王春虎.小麦蛋白组分的连续提取分离法及提取时间的选择[J].河南职业技术师范学院学报,1993,21(2):1-4.]
    [195]李法云,宋丽,官春云,等.辽西半干旱区农田水肥耦合作用对春小麦产量的影响.应用生态学报,2000,11(4),535-539
    [196]吕殿青、张文孝、谷洁等.渭北东部旱塬氮磷水三因素交互作用与耦合模型研究.西北农业学报,1994,3(3)
    [197]于振文,岳松涛,沈成国等.不同密度对冬小麦开花后叶片衰老和粒重的影响.作物学报,1995,21(4),503-508
    [198] Ahmadi A,Baker DA.The effect of water stress on the activities of key regulatory enzymesof the sucrose to starch pathway in wheat.Plant Growth Regul,2001,35:81-91.
    [199] Furbank RT,Foyer CH,Walker DA. Regulation of photosynthesis in isolated spinach chloroplastduring orthophosphate limitation,Biochem,Biophys.Acta,1987,894:552-561.
    [200] Havaux M, Canani O, Malkin S. Photosynthetic responses to water stress in leaves expressedby photoacoustics and related methods Ⅱ The effect of rapid drought on the electrontransport and relative activities of the two photosystems. Plant Physiol,1986,82:834-839.
    [201] Hussain M L, Shan S H, Sajjad H, et al. Growth, yield and quality response of three wheat(Triticum aestivum L.) varieties to different levels of N, P and K[J]. International Journalof Agriculture and Biology,2002,4(3):362–364
    [202] Калинин.气温对小麦生长量的影响.国外农学—麦类作物,1985,(2):33-34.
    [203] Lauer MJ, pallardy SG,Blevirs DG. Whole leaf carbon exchange characteristics of phosphatedeficient soybean,Plant Physiol.,1989,91:848-854.
    [204] Mohanty P, Boyer J S. Chloroplast response to low water poteneials. IV. Quantum yield isreduced. Plant Physiol,1976,57:704-709.
    [205] Xu Xiaoling, Wang Zhimin, Zhang Junping. Effect of heat stress on photosyntheticcharacteristics of different green organs of winter wheat grain-filling stage. ACTA BOTANIASINICA.2001,43(6):571-577.
    [206] Xie Z J,Jiang D. Effects of post-anthesis soil water status on the activities of keyregulatory enzymes of starch and protein accumulation in wheat grains. Journal of PlantPhysiology and Molecular Biology,2003,29(4):309-316.
    [207] Yang J C.Activities of key enzymes in sucrose-to-starch conversion in wheat grain subjectedto water deficit during grain filling.Plant Physiol,2004,135:1621-1629.

© 2004-2018 中国地质图书馆版权所有 京ICP备05064691号 京公网安备11010802017129号

地址:北京市海淀区学院路29号 邮编:100083

电话:办公室:(+86 10)66554848;文献借阅、咨询服务、科技查新:66554700