四川省小麦育成品种系谱分析及发展进程
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摘要
品种系谱蕴含亲本来源、选育方式、亲缘关系等大量信息,对揭示农作物品种演变特点、规律并最终指导育种实践具有重要意义。本文汇总了四川省1936～2017年间326个小麦品种的系谱信息,对育种方式、亲本构成、高频亲本及骨干亲本的变化、遗传贡献、易位系及人工合成种质分布等情况进行了梳理与分析。从20世纪30年代至今,四川省育种家从大量材料中选用387个直接亲本,采用杂交育种等方式,配制数千个组合,从256个组合中选育出314个品种,对四川省小麦生产做出了重要贡献,同时也为小麦育种提供了更为优异的材料基础。四川省小麦育种经历了从无到有,从主要依靠引进材料、地方品种到自主创制材料的过程;高频亲本、骨干亲本随育种进程推进而逐渐变化。易位系和人工合成种质对四川省小麦育种贡献巨大。育种目标的一致性必然导致遗传多样性散失和遗传基础脆弱,今后重点应加强资源的创制、保护和利用。本文通过育成品种的系谱梳理,对四川省小麦的育种发展进程进行了分析,以期为未来种质资源的收集创制和育种研究提供重要参考信息。
Variety pedigree contains a lot of information, including parental origin, breeding methods, genetic relationship, and so on. Studying them could reveal the evolution characteristics and rules of breeding and ultimately guide practice. The pedigrees of 326 wheat varieties from 1936 to 2017 in the history of the Sichuan Province was collected and analyzed in terms of breeding methods, parental composition, changes of high frequency parents and backbone parents,genetic contribution, distribution of translocation lines and synthetic germplasms. Over the past 80 years since 1930s,breeders have selected 387 direct parents from a large number of materials, made 256 combinations by means of cross breeding, and have released 314 varieties from them, which contributed directly to wheat breeding and production in Sichuan. Wheat breeding experienced a process from utilizing landraces, introducing foreign germplasm to creating breeding materials independently; high-frequency parents and backbone parents used for breeding gradually changed in different stage of the breeding history. Synthetic germplasms contributed greatly to wheat breeding in recent years. The consistency of breeding objectives will inevitably lead to the loss of genetic diversity and the fragility of genetic basis. In the future, the protection and utilization of genetic resources should be strengthened. In this review, the development of wheat breeding in Sichuan was summarized through pedigree analysis, in order to provide a reference for future research.
Variety pedigree contains a lot of information, including parental origin, breeding methods, genetic relationship, and so on. Studying them could reveal the evolution characteristics and rules of breeding and ultimately guide practice. The pedigrees of 326 wheat varieties from 1936 to 2017 in the history of the Sichuan Province was collected and analyzed in terms of breeding methods, parental composition, changes of high frequency parents and backbone parents,genetic contribution, distribution of translocation lines and synthetic germplasms. Over the past 80 years since 1930s,breeders have selected 387 direct parents from a large number of materials, made 256 combinations by means of cross breeding, and have released 314 varieties from them, which contributed directly to wheat breeding and production in Sichuan. Wheat breeding experienced a process from utilizing landraces, introducing foreign germplasm to creating breeding materials independently; high-frequency parents and backbone parents used for breeding gradually changed in different stage of the breeding history. Synthetic germplasms contributed greatly to wheat breeding in recent years. The consistency of breeding objectives will inevitably lead to the loss of genetic diversity and the fragility of genetic basis. In the future, the protection and utilization of genetic resources should be strengthened. In this review, the development of wheat breeding in Sichuan was summarized through pedigree analysis, in order to provide a reference for future research.
引文
[1]Jin SB eds.Chinese wheat and its pedigree.Beijing:Agriculture Press,1983,1-417.金善宝主编.中国小麦品种及其系谱.北京:农业出版社,1983,1-417.
[2]Zhuang QS eds.Chinese Wheat Improvement and Pedigree Analysis.Beijing:China Agriculture Press,2003,1-681.庄巧生主编.中国小麦品种改良及系谱分析.北京:中国农业出版社,2003,1-681.
[3]余遥主编.四川小麦.成都:四川科学技术出版社,1998,1-517.
[4]Ren ZL eds.Wheat Breeding in Rainfed Agricultural Areas.Beijing:Science Press,2010,1-288.任正隆主编.雨养农业区的小麦育种.北京:科学出版社,2010,1-288.
[5]Li SR eds.Practice and Development of Wheat Variety Improvement in Mianyang.Beijing:China Agriculture Press,2018,1-261.李生荣主编.绵阳号小麦品种改良的实践与发展.北京:中国农业出版社,2018,1-261.
[6]Zheng JM,Luo JT,Li SZ,Wan HS,Yang MY,Li J,Yang EN,Pu ZJ.Quality of wheat varieties in regional test in Sichuan province from 2008 to 2016.J Trit Crops,2017,37(4):513-519.郑建敏,罗江陶,李式昭,万洪深,杨漫宇,李俊,杨恩年,蒲宗君.2008-2016年四川省小麦区试品系品质分析.麦类作物学报,2017,37(4):513-519.
[7]Zhang ZQ,Zheng YL,Wei YM,Wu W,Zhou YH,Liu DC,Lan XJ.Analysis on genetic diversity among Sichuan wheat cultivars based on SSR markers.J Trit Crops,2002,22(2):5-9.张志清,郑有良,魏育明,吴卫,周永红,刘登才,兰秀锦.四川主栽小麦品种遗传多样性的SSR标记研究.麦类作物学报,2002,22(2):5-9.
[8]Ren ZL,Zhang HQ,Tan FQ,Jiang HR,Yan BJ,Zhuang HY,Fu TH,Zhang XZ,Chen XP.A new variety of wheat,Chuannong 18,with high yield,disease resistance,good quality and early-aging resistance.J Trit Crops,2004,24(4):158.任正隆,张怀琼,谭飞泉,蒋华仁,晏本菊,张怀渝,傅体华,张显志,陈显培.高产抗病优质抗早衰“协调型”小麦新品种--川农18.麦类作物学报,2004,24(4):158.
[9]Li SR.Breeding of large-spike,high-yield and highquality wheat varieties Mianyang 33 and Mianyang 35.Chin Agric Bull,2005,21(4):127-155.李生荣.大穗型丰产优质小麦新品种绵阳33号和绵阳35号的选育.中国农学通报,2005,21(4):127-129,155.
[10]Fu TH,Wang CM,Ren ZL.SSR genetic diversity among modern advanced wheat cultivars(Triticum aestivum L.)in Sichuan and its relationships with their pedigree.JSichuan Agric Univ,2007,25(1):1-8,23.傅体华,王春梅,任正隆.四川育成小麦品种的SSR遗传多态性及系谱关系.四川农业大学学报,2007,25(1):1-8,23.
[11]Wu L,Zhu HZ,Deng L,Hu J.Review of the release wheat cultivars through Sichuan provincial trial during 1997 to2007.Southwest Chin J Agric Sci,2008,21(3):562-569.伍玲,朱华忠,邓丽,胡嘉.1997~2007年通过四川省区试审定的小麦品种述评.西南农业学报,2008,21(3):562-569.
[12]Chen GY,Liu W,He YJ,Gou LL,Yu M,Chen SS,Wei YM,Zheng YL.Specific loci for adult-plant resistance to stripe rust in wheat founder parent Fan 6 and their genetic dissection in its derivatives.Acta Agron Sin,2013,39(5):827-836.陈国跃,刘伟,何员江,苟璐璐,余马,陈时盛,魏育明,郑有良.小麦骨干亲本繁6条锈病成株抗性特异位点及其在衍生品种中的遗传解析.作物学报,2013,39(5):827-836.
[13]Lin ZJ,Jie SH.Review of wheat breeding for 40 years.JHenan Agri Sci,1999,(1):3-4.林作揖,揭声慧.小麦育种工作40年回顾.河南农业科学,1999,(1):3-4.
[14]Fang Z,Zhai DF.A review of winter wheat hybrid breeding in the past sixty years.Shandong Agri Sci,2013,45(3):114-118.方正,翟冬峰.冬小麦杂交育种实践60年回顾.山东农业科学,2013,45(3):114-118.
[15]Zhao TJ,Cui ZL,Gai JY.Nuclear and cytoplasmic contribution of 58-161 to the released soybean cultivars in China.Soyb Sci,1998,17(2):120-128.赵团结,崔章林,盖钧镒.中国大豆育成品种中江苏种质58-161的遗传贡献.大豆科学,1998,17(2):120-128.
[16]Gai JY,Zhao TJ,Cui ZL,Qiu JX.Nuclear and cytoplasmic contributions of germplasm from distinct areas to the soybean cultivars released during 1923-1995in China.Sci Agric Sin,1998,31(5):35-43.盖钧镒,赵团结,崔章林,邱家驯.中国大豆育成品种中不同地理来源种质的遗传贡献.中国农业科学,1998,31(5):35-43.
[17]He ZH,Zhuang QS,Cheng SH,Yu ZW,Zhao ZD,Liu X.Wheat production and technology improvement in China.J Agric,2018,8(1):107-114.何中虎,庄巧生,程顺和,于振文,赵振东,刘旭.中国小麦产业发展与科技进步.农学学报,2018,8(1):107-114.
[18]Zhang XY,Tong YP,You GX,Hao CY,Gai HM,Wang LF,Li B,Dong YC,Li ZS.Hitchhiking effect mapping:a new approach for discovering agronomic important genes.Sci Agric Sin,2006,39(8):1526-1535.张学勇,童依平,游光霞,郝晨阳,盖红梅,王兰芬,李滨,董玉琛,李振声.选择牵连效应分析:发掘重要基因的新思路.中国农业科学,2006,39(8):1526-1535.
[19]Qi YW,Zhang DL,Zhang HL,Wang MX,Sun JL,Liao DQ,Wei XH,Qiu ZE,Tang SX,Cao YS,Wang XK,Li ZC.Genetic diversity of rice breeding varieties in China and its variation trend in recent 50 years.Chin Sci Bull,2006,51(6):693-699.齐永文,张冬玲,张洪亮,王美兴,孙俊立,廖登群,魏兴华,裘宗恩,汤圣祥,曹永生,王象坤,李自超.中国水稻选育品种遗传多样性及其近50年变化趋势.科学通报,2006,51(6):693-699.
[20]Hua L,Yuan XP,Yu HY,Wang YP,Xu Q,Tang SX,Wei XH.A comparative study on SSR diversity in Chinese major rice varieties planted in 1950s and during the most recent ten years.Chin J Rice Sci,2007,21(2):150-154.华蕾,袁筱萍,余汉勇,王一平,徐群,汤圣祥,魏兴华.我国水稻主栽品种SSR多样性的比较分析.中国水稻科学,2007,21(2):150-154.
[21]Zhang SH,Tian QZ,Li XH,Li MS,Xie CX.Advancement of maize germplasm improvement and relevant research.JMaize Sci,2006,14(1):1-6.张世煌,田清震,李新海,李明顺,谢传晓.玉米种质改良与相关理论研究进展.玉米科学,2006,14(1):1-6.
[22]Meng YJ,Yan JB,Teng WT,Li JS.Trends in genetic diversity among widely used inbreds from 1991 to 2001 in China and application of three major germplasm groups in maize breeding.Sci Agric Sin,2010,43(4):670-679.孟义江,严建兵,滕文涛,李建生.1991-2001年中国主要玉米杂交种遗传基础的变化趋势及三大种质类群在育种中的应用.中国农业科学,2010,43(4):670-679.
[23]Jia JZ,Zhang ZB,Devos K,Gale MD.Genetic diversity analysis of RFLP mapping sites on 21 chromosomes in wheat.Science in China(Series C:Life Sciences),2001,31(1):13-21.贾继增,张正斌,Devos K,Gale MD.小麦21条染色体RFLP作图位点遗传多样性分析.中国科学C辑,2001,31(1):13-21.
[24]Hao CY,Wang LF,Zhang XY,You GX,Dong YC,Jia JZ,Liu X,Shan XW,Liu SC,Cao YS.Evolution of genetic diversity of cultivated wheat varieties in China.Science in China(Series C:Life Sciences),2005,35(5):408-415.郝晨阳,王兰芬,张学勇,游光霞,董玉琛,贾继增,刘旭,尚勋武,刘三才,曹永生.我国育成小麦品种的遗传多样性演变.中国科学C辑,2005,35(5):408-415.
[25]Duan YF,Liu J,Bing CS,Duan SG,Xu JF,Jin LP.Construction of fingerprinting and analysis of genetic diversity with SSR markers for eighty-eight approved potato cultivars(Solanum tuberosum L.)in China.Acta Agron Sin,2009,35(8):1451-1457.段艳凤,刘杰,卞春松,段绍光,徐建飞,金黎平.中国88个马铃薯审定品种SSR指纹图谱构建与遗传多样性分析.作物学报,2009,35(8):1451-1457.
[26]Wang HG,Jia GQ,Zhi H,Wen QF,Dong JL,Chen L,Wang JJ,Cao XN,Liu SC,Wang L,Qiao ZJ,Diao XM.Phenotypic diversity evaluations of foxtail millet core collections.Acta Agron Sin,2016,42(1):19-30.王海岗,贾冠清,智慧,温琪汾,董俊丽,陈凌,王君杰,曹晓宁,刘思辰,王纶,乔治军,刁现民.谷子核心种质表型遗传多样性分析及综合评价.作物学报,2016,42(1):19-30.
[27]Li Q,Liu QC,Zhai H,Ma DF,Wang X,Li XQ,Wang YP.Genetic diversity in main parents of sweetpotato in China as revealed by ISSR Marker.Acta Agron Sin,2008,34(6):972-977.李强,刘庆昌,翟红,马代夫,王欣,李雪琴,王玉萍.中国甘薯主要亲本遗传多样性的ISSR分析.作物学报,2008,34(6):972-977.
[28]Lai Y,Wang PX,Fan GQ,Si EJ,Wang J,Yang K,Meng YX,Li BC,Ma XL,Shang XW,Wang HJ.Genetic diversity and association analysis using SSR markers in barley.Sci Agric Sin,2013,46(2):233-242.赖勇,王鹏喜,范贵强,司二静,王晋,杨轲,孟亚雄,李葆春,马小乐,尚勋武,王化俊.大麦SSR标记遗传多样性及其与农艺性状关联分析.中国农业科学,2013,46(2):233-242.
[29]Sun ZQ,Qi FY,Zheng Z,Dong WZ,Huang BY,Zhang J,Zhang ZX,Tang FS,Zhang XY,Liu ZY.Analysis of parental relationship of 106 peanut cultivars released in Henan province.Chin J Oil Crop Sci,2017,39(6):754-762.孙子淇,齐飞艳,郑峥,董文召,黄冰艳,张俊,张忠信,汤丰收,张新友,刘志勇.河南省106个审定花生品种亲缘关系分析.中国油料作物学报,2017,39(6):754-762.
[30]Wang ZT,You Q,Gao SW,Wang CF,Li Z,Ma JJ,Que YX,Xu LP,Luo J.Identification of sugarcane varieties by AFLP and SSR markers and its application.Acta Agron Sin,2018,44(5):723-736.汪洲涛,游倩,高世武,王春风,李竹,马晶晶,阙友雄,许莉萍,罗俊.甘蔗品种的AFLP和SSR标记鉴定及其应用.作物学报,2018,44(5):723-736.
[31]Shen L,Hua YF,Fu YP,Li J,Liu Q,Jiao XZ,Xin GW,Wang JJ,Wang XC,Yan CJ,Wang KJ.Rapid generation of genetic diversity by multiplex CRISPR/Cas9 genome editing in rice.Sci Chin-Ear Sci,2017,60:506-515.沈兰,华宇峰,付亚萍,李健,刘庆,焦晓真,辛高伟,王俊杰,王兴春,严长杰,王克剑.利用CRISPR/Cas9多基因编辑系统在水稻中快速引入遗传多样性.中国科学,2017,47(11):1186-1195.
[32]Zhou XC,Xing YZ.The application of genome editing in identification of plant gene function and crop breeding.Hereditas(Beijing),2016,38(3):227-242.周想春,邢永忠.基因组编辑技术在植物基因功能鉴定及作物育种中的应用.遗传,2016,38(3):227-242.
[33]An DG,Xu HX,Xu YF.Enhancement of wheat distant hybridization germplasm.Chin J Eco-Agric,2011,19(5):1011-1019.安调过,许红星,许云峰.小麦远缘杂交种质资源创新.中国生态农业学报,2011,19(5):1011-1019.
[34]Wang LH,Zhou M,Li HL,He ZH,Xia XC.Cloning and phylogenetic analysis of low-molecular-weight glutenin subunit genes at Glu-B3 locus in common wheat relative species.Hereditas(Beijing),2010,32(6):613-624.王林海,周敏,李慧玲,何中虎,夏先春.小麦近缘种低分子量麦谷蛋白亚基基因Glu-B3克隆及系统发育分析.遗传,2010,32(6):613-624.
[35]Li J,Zhu XG,Wan HS,Wang Q,Tang ZX,Fu SL,Yang ZJ,Yang MY,Yang WY.Identification of the 1RS-7DS.7DLwheat-rye small segment translocation lines.Hereditas(Beijing),2015,37(6):590-598.李俊,朱欣果,万洪深,王琴,唐宗祥,符书兰,杨足君,杨漫宇,杨武云.1RS-7DS.7DL小麦-黑麦小片段易位系的鉴定.遗传,2015,37(6):590-598.
[36]Rabinovich SV.Importance of wheat-rye translocations for breeding modern cultivars of Triticum aestivum L.Euphytica,1998,100:323-340.
[37]Zhou Y,He ZH,Zhang GS,Xia LQ,Chen XM,Gao YC,Jing ZB,Yu GJ.Utilization of 1BL/1RS translocation in wheat breeding in China.Acta Agron Sin,2004,30(4):531-535.周阳,何中虎,张改生,夏兰琴,陈新民,高永超,井赵斌,于广军.1BL/1RS易位系在我国小麦育种中的应用.作物学报,2004,30(6):531-535.
[38]Ren TH,Chen F,Zhuang HQ,Yan BJ,Ren ZL.Application of 1RS.1BL translocation in the breeding of“Chuannong”series wheat cuitivars.J Trit Crops,2011,31(3):430-436.任天恒,陈放,张怀琼,晏本菊,任正隆.1RS.1BL易位在川农号系列小麦新品种选育中的作用.麦类作物学报,2011,31(3):430-436.
[39]Ren ZL,Zhang HQ.Induction of translocation of small fragments of wheat-rye chromosomes.Science in China(Series C:Life Sciences),1997,27(3):258-263.任正隆,张怀琼.小麦-黑麦染色体小片段易位的诱导.中国科学C辑,1997,27(3):258-263.
[40]Ren TH,Yan BJ,Zhang HQ,Ren ZL.Effect of 1RS.1BLtranslocation chromosome on stay green trait in common wheat(Triticum aestivun L.).J Trit Crops,2009,29(3):419-423.任天恒,晏本菊,张怀琼,任正隆.1RS.1BL易位染色体对小麦开花后叶片延绿特性的影响.麦类作物学报,2009,29(3):419-423.
[41]Ren TH,Li Z,Yan BJ,Yang MY,Tan FQ,Ren ZL.Development of a novel primary 1RS.1BL translocation line T956-13 and its breeding value.J Trit Crops,2017,37(12):1534-1540.任天恒,李治,晏本菊,杨漫宇,谭飞泉,任正隆.新育成的1RS·1BL初级易位系T956-13的育种价值.麦类作物学报,2017,37(12):1534-1540.
[42]Liu DJ,Qi LL,Chen PD,Zhou B,Zhang SZ.Precise identification of alien chromosome segment introduced in wheat and the stability of its resistance gene.Acta Genet Sin,1996,23(1):18-23.刘大均,齐莉莉,陈佩度,周波,张守中.导入小麦的外源染色体片段的准确鉴定及外源抗性基因的稳定性分析.遗传学报,1996,23(1):18-23.
[43]Qi LL,Chen PD,Liu DJ,Zhou B,Zhang SZ,Sheng BQ,Xiang QJ,Duang XY,Zhou YL.The gene Pm21-a new source for resistance to wheat powdery mildew.Acta Agron Sin,1995,21(3):257-262.齐莉莉,陈佩度,刘大均,周波,张守中,盛宝钦,向齐君,段霞渝,周益林.小麦白粉病新抗源-基因Pm21.作物学报,1995,21(3):257-262.
[44]Chen P,Qi L,Zhou B,Zhang S,Liu D.Development and molecular cytogenetic analysis of wheat-Haynaldia villosa6VS/6AL translocation lines specifying resistance to powdery mildew.Theor Appl Genet,1995,91(6-7):1125-1128.
[45]Li G,Chen P,Zhang S,Wang X,He Z,Zhang Y,Zhao H,Huang H,Zhou X.Effects of the 6VS.6AL translocation on agronomic traits and dough properties of wheat.Euphytica,2008,155(3):305-313.
[46]Yang W,Liu D,Li J,Zhang L,Wei H,Hu X,Zheng Y,He ZH,Zou Y.Synthetic hexaploid wheat and its utilization for wheat genetic improvement in China.J Genet Genomics,2009,36(9):539-546.
[47]Li J,Wei HT,Yang SJ,Li CS,Tang YL,Hu XR,Yang WY.Genetic effects of 1BS chromosome arm on the main agronomic traits in Chuanmai 42.Acta Agron Sin,2009,35(12):2167-2173.李俊,魏会廷,杨粟洁,李朝苏,汤永禄,胡晓蓉,杨武云.川麦42的1BS染色体臂对小麦主要农艺性状的遗传效应.作物学报,2009,35(12):2167-2173.
[48]Li J,Wei HT,Hu XR,Li CS,Tang YL,Liu DC,Yang WY.Identification of a high-yield introgression locus from synthetic hexaploid wheat in Chuanmai 42.Acta Agron Sin,2011,37(2):255-262.李俊,魏会廷,胡晓蓉,李朝苏,汤永禄,刘登才,杨武云.川麦42中源于人工合成小麦的一个高产位点鉴定.作物学报,2011,37(2):255-262.
[49]Li J,Wei HT,Yang SJ,Li CS,Tang YL,Hu XR,Yang WY.Genetic effects of 1BS chromosome arm on the main agronomic traits in Chuanmai 42.Acta Agron Sin,2009,35(12):2167-2173.李俊,魏会廷,杨粟洁,李朝苏,汤永禄,胡晓蓉,杨武云.川麦42的1BS染色体臂对小麦主要农艺性状的遗传效应.作物学报,2009,35(12):2167-2173.
[50]Wan H,Yang Y,Li J,Zhang Z,Yang W.Mapping a major QTL for hairy leaf sheath introgressed from Aegilops tauschii and its association with enhanced grain yield in bread wheat.Euphytica,2015,205(1):275-285.
[51]Li J,Wan HS,Yang WY,Wang Q,Zhu XG,Hu XR,Wei HT,Tang YL,Li CS,Peng ZS,Zhou YH.Dissection of genetic components in the new high-yielding wheat cultivar Chuanmai 104.Sci Agric Sin,2014,47:2281-2291.李俊,万洪深,杨武云,王琴,朱欣果,胡晓蓉,魏会廷,汤永禄,李朝苏,彭正松,周永红.小麦新品种川麦104的遗传构成分析.中国农业科学,2014,47(12):2281-2291.
[52]Liu DC,Zhang LQ,Hao M,Ning SZ,Yuan ZW,Dai SF,Huang L,Wu BH,Yan ZH,Lan XJ,Zheng YL.Wheat breeding in the hometown of Chinese Spring.The Crop J,2018,6(1):82-90.
[2]Zhuang QS eds.Chinese Wheat Improvement and Pedigree Analysis.Beijing:China Agriculture Press,2003,1-681.庄巧生主编.中国小麦品种改良及系谱分析.北京:中国农业出版社,2003,1-681.
[3]余遥主编.四川小麦.成都:四川科学技术出版社,1998,1-517.
[4]Ren ZL eds.Wheat Breeding in Rainfed Agricultural Areas.Beijing:Science Press,2010,1-288.任正隆主编.雨养农业区的小麦育种.北京:科学出版社,2010,1-288.
[5]Li SR eds.Practice and Development of Wheat Variety Improvement in Mianyang.Beijing:China Agriculture Press,2018,1-261.李生荣主编.绵阳号小麦品种改良的实践与发展.北京:中国农业出版社,2018,1-261.
[6]Zheng JM,Luo JT,Li SZ,Wan HS,Yang MY,Li J,Yang EN,Pu ZJ.Quality of wheat varieties in regional test in Sichuan province from 2008 to 2016.J Trit Crops,2017,37(4):513-519.郑建敏,罗江陶,李式昭,万洪深,杨漫宇,李俊,杨恩年,蒲宗君.2008-2016年四川省小麦区试品系品质分析.麦类作物学报,2017,37(4):513-519.
[7]Zhang ZQ,Zheng YL,Wei YM,Wu W,Zhou YH,Liu DC,Lan XJ.Analysis on genetic diversity among Sichuan wheat cultivars based on SSR markers.J Trit Crops,2002,22(2):5-9.张志清,郑有良,魏育明,吴卫,周永红,刘登才,兰秀锦.四川主栽小麦品种遗传多样性的SSR标记研究.麦类作物学报,2002,22(2):5-9.
[8]Ren ZL,Zhang HQ,Tan FQ,Jiang HR,Yan BJ,Zhuang HY,Fu TH,Zhang XZ,Chen XP.A new variety of wheat,Chuannong 18,with high yield,disease resistance,good quality and early-aging resistance.J Trit Crops,2004,24(4):158.任正隆,张怀琼,谭飞泉,蒋华仁,晏本菊,张怀渝,傅体华,张显志,陈显培.高产抗病优质抗早衰“协调型”小麦新品种--川农18.麦类作物学报,2004,24(4):158.
[9]Li SR.Breeding of large-spike,high-yield and highquality wheat varieties Mianyang 33 and Mianyang 35.Chin Agric Bull,2005,21(4):127-155.李生荣.大穗型丰产优质小麦新品种绵阳33号和绵阳35号的选育.中国农学通报,2005,21(4):127-129,155.
[10]Fu TH,Wang CM,Ren ZL.SSR genetic diversity among modern advanced wheat cultivars(Triticum aestivum L.)in Sichuan and its relationships with their pedigree.JSichuan Agric Univ,2007,25(1):1-8,23.傅体华,王春梅,任正隆.四川育成小麦品种的SSR遗传多态性及系谱关系.四川农业大学学报,2007,25(1):1-8,23.
[11]Wu L,Zhu HZ,Deng L,Hu J.Review of the release wheat cultivars through Sichuan provincial trial during 1997 to2007.Southwest Chin J Agric Sci,2008,21(3):562-569.伍玲,朱华忠,邓丽,胡嘉.1997~2007年通过四川省区试审定的小麦品种述评.西南农业学报,2008,21(3):562-569.
[12]Chen GY,Liu W,He YJ,Gou LL,Yu M,Chen SS,Wei YM,Zheng YL.Specific loci for adult-plant resistance to stripe rust in wheat founder parent Fan 6 and their genetic dissection in its derivatives.Acta Agron Sin,2013,39(5):827-836.陈国跃,刘伟,何员江,苟璐璐,余马,陈时盛,魏育明,郑有良.小麦骨干亲本繁6条锈病成株抗性特异位点及其在衍生品种中的遗传解析.作物学报,2013,39(5):827-836.
[13]Lin ZJ,Jie SH.Review of wheat breeding for 40 years.JHenan Agri Sci,1999,(1):3-4.林作揖,揭声慧.小麦育种工作40年回顾.河南农业科学,1999,(1):3-4.
[14]Fang Z,Zhai DF.A review of winter wheat hybrid breeding in the past sixty years.Shandong Agri Sci,2013,45(3):114-118.方正,翟冬峰.冬小麦杂交育种实践60年回顾.山东农业科学,2013,45(3):114-118.
[15]Zhao TJ,Cui ZL,Gai JY.Nuclear and cytoplasmic contribution of 58-161 to the released soybean cultivars in China.Soyb Sci,1998,17(2):120-128.赵团结,崔章林,盖钧镒.中国大豆育成品种中江苏种质58-161的遗传贡献.大豆科学,1998,17(2):120-128.
[16]Gai JY,Zhao TJ,Cui ZL,Qiu JX.Nuclear and cytoplasmic contributions of germplasm from distinct areas to the soybean cultivars released during 1923-1995in China.Sci Agric Sin,1998,31(5):35-43.盖钧镒,赵团结,崔章林,邱家驯.中国大豆育成品种中不同地理来源种质的遗传贡献.中国农业科学,1998,31(5):35-43.
[17]He ZH,Zhuang QS,Cheng SH,Yu ZW,Zhao ZD,Liu X.Wheat production and technology improvement in China.J Agric,2018,8(1):107-114.何中虎,庄巧生,程顺和,于振文,赵振东,刘旭.中国小麦产业发展与科技进步.农学学报,2018,8(1):107-114.
[18]Zhang XY,Tong YP,You GX,Hao CY,Gai HM,Wang LF,Li B,Dong YC,Li ZS.Hitchhiking effect mapping:a new approach for discovering agronomic important genes.Sci Agric Sin,2006,39(8):1526-1535.张学勇,童依平,游光霞,郝晨阳,盖红梅,王兰芬,李滨,董玉琛,李振声.选择牵连效应分析:发掘重要基因的新思路.中国农业科学,2006,39(8):1526-1535.
[19]Qi YW,Zhang DL,Zhang HL,Wang MX,Sun JL,Liao DQ,Wei XH,Qiu ZE,Tang SX,Cao YS,Wang XK,Li ZC.Genetic diversity of rice breeding varieties in China and its variation trend in recent 50 years.Chin Sci Bull,2006,51(6):693-699.齐永文,张冬玲,张洪亮,王美兴,孙俊立,廖登群,魏兴华,裘宗恩,汤圣祥,曹永生,王象坤,李自超.中国水稻选育品种遗传多样性及其近50年变化趋势.科学通报,2006,51(6):693-699.
[20]Hua L,Yuan XP,Yu HY,Wang YP,Xu Q,Tang SX,Wei XH.A comparative study on SSR diversity in Chinese major rice varieties planted in 1950s and during the most recent ten years.Chin J Rice Sci,2007,21(2):150-154.华蕾,袁筱萍,余汉勇,王一平,徐群,汤圣祥,魏兴华.我国水稻主栽品种SSR多样性的比较分析.中国水稻科学,2007,21(2):150-154.
[21]Zhang SH,Tian QZ,Li XH,Li MS,Xie CX.Advancement of maize germplasm improvement and relevant research.JMaize Sci,2006,14(1):1-6.张世煌,田清震,李新海,李明顺,谢传晓.玉米种质改良与相关理论研究进展.玉米科学,2006,14(1):1-6.
[22]Meng YJ,Yan JB,Teng WT,Li JS.Trends in genetic diversity among widely used inbreds from 1991 to 2001 in China and application of three major germplasm groups in maize breeding.Sci Agric Sin,2010,43(4):670-679.孟义江,严建兵,滕文涛,李建生.1991-2001年中国主要玉米杂交种遗传基础的变化趋势及三大种质类群在育种中的应用.中国农业科学,2010,43(4):670-679.
[23]Jia JZ,Zhang ZB,Devos K,Gale MD.Genetic diversity analysis of RFLP mapping sites on 21 chromosomes in wheat.Science in China(Series C:Life Sciences),2001,31(1):13-21.贾继增,张正斌,Devos K,Gale MD.小麦21条染色体RFLP作图位点遗传多样性分析.中国科学C辑,2001,31(1):13-21.
[24]Hao CY,Wang LF,Zhang XY,You GX,Dong YC,Jia JZ,Liu X,Shan XW,Liu SC,Cao YS.Evolution of genetic diversity of cultivated wheat varieties in China.Science in China(Series C:Life Sciences),2005,35(5):408-415.郝晨阳,王兰芬,张学勇,游光霞,董玉琛,贾继增,刘旭,尚勋武,刘三才,曹永生.我国育成小麦品种的遗传多样性演变.中国科学C辑,2005,35(5):408-415.
[25]Duan YF,Liu J,Bing CS,Duan SG,Xu JF,Jin LP.Construction of fingerprinting and analysis of genetic diversity with SSR markers for eighty-eight approved potato cultivars(Solanum tuberosum L.)in China.Acta Agron Sin,2009,35(8):1451-1457.段艳凤,刘杰,卞春松,段绍光,徐建飞,金黎平.中国88个马铃薯审定品种SSR指纹图谱构建与遗传多样性分析.作物学报,2009,35(8):1451-1457.
[26]Wang HG,Jia GQ,Zhi H,Wen QF,Dong JL,Chen L,Wang JJ,Cao XN,Liu SC,Wang L,Qiao ZJ,Diao XM.Phenotypic diversity evaluations of foxtail millet core collections.Acta Agron Sin,2016,42(1):19-30.王海岗,贾冠清,智慧,温琪汾,董俊丽,陈凌,王君杰,曹晓宁,刘思辰,王纶,乔治军,刁现民.谷子核心种质表型遗传多样性分析及综合评价.作物学报,2016,42(1):19-30.
[27]Li Q,Liu QC,Zhai H,Ma DF,Wang X,Li XQ,Wang YP.Genetic diversity in main parents of sweetpotato in China as revealed by ISSR Marker.Acta Agron Sin,2008,34(6):972-977.李强,刘庆昌,翟红,马代夫,王欣,李雪琴,王玉萍.中国甘薯主要亲本遗传多样性的ISSR分析.作物学报,2008,34(6):972-977.
[28]Lai Y,Wang PX,Fan GQ,Si EJ,Wang J,Yang K,Meng YX,Li BC,Ma XL,Shang XW,Wang HJ.Genetic diversity and association analysis using SSR markers in barley.Sci Agric Sin,2013,46(2):233-242.赖勇,王鹏喜,范贵强,司二静,王晋,杨轲,孟亚雄,李葆春,马小乐,尚勋武,王化俊.大麦SSR标记遗传多样性及其与农艺性状关联分析.中国农业科学,2013,46(2):233-242.
[29]Sun ZQ,Qi FY,Zheng Z,Dong WZ,Huang BY,Zhang J,Zhang ZX,Tang FS,Zhang XY,Liu ZY.Analysis of parental relationship of 106 peanut cultivars released in Henan province.Chin J Oil Crop Sci,2017,39(6):754-762.孙子淇,齐飞艳,郑峥,董文召,黄冰艳,张俊,张忠信,汤丰收,张新友,刘志勇.河南省106个审定花生品种亲缘关系分析.中国油料作物学报,2017,39(6):754-762.
[30]Wang ZT,You Q,Gao SW,Wang CF,Li Z,Ma JJ,Que YX,Xu LP,Luo J.Identification of sugarcane varieties by AFLP and SSR markers and its application.Acta Agron Sin,2018,44(5):723-736.汪洲涛,游倩,高世武,王春风,李竹,马晶晶,阙友雄,许莉萍,罗俊.甘蔗品种的AFLP和SSR标记鉴定及其应用.作物学报,2018,44(5):723-736.
[31]Shen L,Hua YF,Fu YP,Li J,Liu Q,Jiao XZ,Xin GW,Wang JJ,Wang XC,Yan CJ,Wang KJ.Rapid generation of genetic diversity by multiplex CRISPR/Cas9 genome editing in rice.Sci Chin-Ear Sci,2017,60:506-515.沈兰,华宇峰,付亚萍,李健,刘庆,焦晓真,辛高伟,王俊杰,王兴春,严长杰,王克剑.利用CRISPR/Cas9多基因编辑系统在水稻中快速引入遗传多样性.中国科学,2017,47(11):1186-1195.
[32]Zhou XC,Xing YZ.The application of genome editing in identification of plant gene function and crop breeding.Hereditas(Beijing),2016,38(3):227-242.周想春,邢永忠.基因组编辑技术在植物基因功能鉴定及作物育种中的应用.遗传,2016,38(3):227-242.
[33]An DG,Xu HX,Xu YF.Enhancement of wheat distant hybridization germplasm.Chin J Eco-Agric,2011,19(5):1011-1019.安调过,许红星,许云峰.小麦远缘杂交种质资源创新.中国生态农业学报,2011,19(5):1011-1019.
[34]Wang LH,Zhou M,Li HL,He ZH,Xia XC.Cloning and phylogenetic analysis of low-molecular-weight glutenin subunit genes at Glu-B3 locus in common wheat relative species.Hereditas(Beijing),2010,32(6):613-624.王林海,周敏,李慧玲,何中虎,夏先春.小麦近缘种低分子量麦谷蛋白亚基基因Glu-B3克隆及系统发育分析.遗传,2010,32(6):613-624.
[35]Li J,Zhu XG,Wan HS,Wang Q,Tang ZX,Fu SL,Yang ZJ,Yang MY,Yang WY.Identification of the 1RS-7DS.7DLwheat-rye small segment translocation lines.Hereditas(Beijing),2015,37(6):590-598.李俊,朱欣果,万洪深,王琴,唐宗祥,符书兰,杨足君,杨漫宇,杨武云.1RS-7DS.7DL小麦-黑麦小片段易位系的鉴定.遗传,2015,37(6):590-598.
[36]Rabinovich SV.Importance of wheat-rye translocations for breeding modern cultivars of Triticum aestivum L.Euphytica,1998,100:323-340.
[37]Zhou Y,He ZH,Zhang GS,Xia LQ,Chen XM,Gao YC,Jing ZB,Yu GJ.Utilization of 1BL/1RS translocation in wheat breeding in China.Acta Agron Sin,2004,30(4):531-535.周阳,何中虎,张改生,夏兰琴,陈新民,高永超,井赵斌,于广军.1BL/1RS易位系在我国小麦育种中的应用.作物学报,2004,30(6):531-535.
[38]Ren TH,Chen F,Zhuang HQ,Yan BJ,Ren ZL.Application of 1RS.1BL translocation in the breeding of“Chuannong”series wheat cuitivars.J Trit Crops,2011,31(3):430-436.任天恒,陈放,张怀琼,晏本菊,任正隆.1RS.1BL易位在川农号系列小麦新品种选育中的作用.麦类作物学报,2011,31(3):430-436.
[39]Ren ZL,Zhang HQ.Induction of translocation of small fragments of wheat-rye chromosomes.Science in China(Series C:Life Sciences),1997,27(3):258-263.任正隆,张怀琼.小麦-黑麦染色体小片段易位的诱导.中国科学C辑,1997,27(3):258-263.
[40]Ren TH,Yan BJ,Zhang HQ,Ren ZL.Effect of 1RS.1BLtranslocation chromosome on stay green trait in common wheat(Triticum aestivun L.).J Trit Crops,2009,29(3):419-423.任天恒,晏本菊,张怀琼,任正隆.1RS.1BL易位染色体对小麦开花后叶片延绿特性的影响.麦类作物学报,2009,29(3):419-423.
[41]Ren TH,Li Z,Yan BJ,Yang MY,Tan FQ,Ren ZL.Development of a novel primary 1RS.1BL translocation line T956-13 and its breeding value.J Trit Crops,2017,37(12):1534-1540.任天恒,李治,晏本菊,杨漫宇,谭飞泉,任正隆.新育成的1RS·1BL初级易位系T956-13的育种价值.麦类作物学报,2017,37(12):1534-1540.
[42]Liu DJ,Qi LL,Chen PD,Zhou B,Zhang SZ.Precise identification of alien chromosome segment introduced in wheat and the stability of its resistance gene.Acta Genet Sin,1996,23(1):18-23.刘大均,齐莉莉,陈佩度,周波,张守中.导入小麦的外源染色体片段的准确鉴定及外源抗性基因的稳定性分析.遗传学报,1996,23(1):18-23.
[43]Qi LL,Chen PD,Liu DJ,Zhou B,Zhang SZ,Sheng BQ,Xiang QJ,Duang XY,Zhou YL.The gene Pm21-a new source for resistance to wheat powdery mildew.Acta Agron Sin,1995,21(3):257-262.齐莉莉,陈佩度,刘大均,周波,张守中,盛宝钦,向齐君,段霞渝,周益林.小麦白粉病新抗源-基因Pm21.作物学报,1995,21(3):257-262.
[44]Chen P,Qi L,Zhou B,Zhang S,Liu D.Development and molecular cytogenetic analysis of wheat-Haynaldia villosa6VS/6AL translocation lines specifying resistance to powdery mildew.Theor Appl Genet,1995,91(6-7):1125-1128.
[45]Li G,Chen P,Zhang S,Wang X,He Z,Zhang Y,Zhao H,Huang H,Zhou X.Effects of the 6VS.6AL translocation on agronomic traits and dough properties of wheat.Euphytica,2008,155(3):305-313.
[46]Yang W,Liu D,Li J,Zhang L,Wei H,Hu X,Zheng Y,He ZH,Zou Y.Synthetic hexaploid wheat and its utilization for wheat genetic improvement in China.J Genet Genomics,2009,36(9):539-546.
[47]Li J,Wei HT,Yang SJ,Li CS,Tang YL,Hu XR,Yang WY.Genetic effects of 1BS chromosome arm on the main agronomic traits in Chuanmai 42.Acta Agron Sin,2009,35(12):2167-2173.李俊,魏会廷,杨粟洁,李朝苏,汤永禄,胡晓蓉,杨武云.川麦42的1BS染色体臂对小麦主要农艺性状的遗传效应.作物学报,2009,35(12):2167-2173.
[48]Li J,Wei HT,Hu XR,Li CS,Tang YL,Liu DC,Yang WY.Identification of a high-yield introgression locus from synthetic hexaploid wheat in Chuanmai 42.Acta Agron Sin,2011,37(2):255-262.李俊,魏会廷,胡晓蓉,李朝苏,汤永禄,刘登才,杨武云.川麦42中源于人工合成小麦的一个高产位点鉴定.作物学报,2011,37(2):255-262.
[49]Li J,Wei HT,Yang SJ,Li CS,Tang YL,Hu XR,Yang WY.Genetic effects of 1BS chromosome arm on the main agronomic traits in Chuanmai 42.Acta Agron Sin,2009,35(12):2167-2173.李俊,魏会廷,杨粟洁,李朝苏,汤永禄,胡晓蓉,杨武云.川麦42的1BS染色体臂对小麦主要农艺性状的遗传效应.作物学报,2009,35(12):2167-2173.
[50]Wan H,Yang Y,Li J,Zhang Z,Yang W.Mapping a major QTL for hairy leaf sheath introgressed from Aegilops tauschii and its association with enhanced grain yield in bread wheat.Euphytica,2015,205(1):275-285.
[51]Li J,Wan HS,Yang WY,Wang Q,Zhu XG,Hu XR,Wei HT,Tang YL,Li CS,Peng ZS,Zhou YH.Dissection of genetic components in the new high-yielding wheat cultivar Chuanmai 104.Sci Agric Sin,2014,47:2281-2291.李俊,万洪深,杨武云,王琴,朱欣果,胡晓蓉,魏会廷,汤永禄,李朝苏,彭正松,周永红.小麦新品种川麦104的遗传构成分析.中国农业科学,2014,47(12):2281-2291.
[52]Liu DC,Zhang LQ,Hao M,Ning SZ,Yuan ZW,Dai SF,Huang L,Wu BH,Yan ZH,Lan XJ,Zheng YL.Wheat breeding in the hometown of Chinese Spring.The Crop J,2018,6(1):82-90.
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