小麦株型建成及氮素调控研究
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摘要
良好的小麦株型是获得高产的重要基础,因此,研究并构建小麦高效冠层的调控途径,具有重要的理论意义和应用价值。本研究以不同株型特征的小麦品种(紧凑型高秆宁麦9号、紧凑型矮秆矮抗58、中间型扬麦12和松散型淮麦17)为对象,研究不同施氮水平下,小麦叶型、茎型和群体冠层结构特征的变化规律,以及株型特征与籽粒产量和构成因素之间的关系,并最终提出理想株型的调控途径。
本研究以不同氮素水平下、不同株型小麦品种的两年田间试验为基础,通过连续观测不同处理条件下小麦主茎叶片形态指标(包括叶长、叶宽、茎叶夹角、叶间距等),综合分析了小麦叶片形态指标随生育进程的变化规律,以及氮素和株型对叶片形态的调控效应。结果表明,随着生长度日的逐渐增加,小麦主茎叶片长度和宽度、茎叶夹角、单株叶面积均呈“S”形曲线变化,而叶片SPAD值,至开花期达最高值,随后逐渐下降。随着叶位的增加,小麦叶长和叶宽、茎叶夹角均符合二次多项式曲线,而单叶面积呈“升-降-升-降”的变化趋势。不同株型小麦品种顶部三张叶片的叶型指标差异主要表现在顶部三张叶片的叶长和SPAD值,旗叶与倒二叶间距及倒三叶基角。施氮量对不同株型小麦品种叶型指标的建成过程有显著影响,但对不同品种的影响程度有差异。
通过连续观测不同处理条件下小麦株高和主茎节间构成,综合分析了小麦株高及其构成、节间直径和长度随生育进程的变化规律,及氮素和株型对其的调控效应。结果表明,小麦株高随生长度日的动态伸长过程可以采用Logistic方程进行描述。小麦主茎顶四节的节间长大小顺序为：穗下节间>倒二节间>倒三节间>倒四节间,且均以穗下节间最细；紧凑型宁麦9号和矮抗58以倒四节间最粗,而松散型淮麦17和中间型扬麦12以倒二节间最粗。株高拟合方程的参数及衍生参数在不同施氮量和株型间存在显著差异。小麦主茎顶四节间的节间长和节间粗、穗长与施氮量呈正相关。在拔节、齐穗、灌浆和成熟期,施氮量对株高均有显著影响,而在返青期对株高的影响不显著。于齐穗、灌浆和成熟期,施氮量和株型对节间长和直径有显著影响,但对株高构成指数影响不显著。
基于不同氮素水平下、不同株型小麦品种叶面积指数、叶面积密度分布、透光率和光合速率等冠层特征的连续观测数据,综合分析了小麦冠层结构特征随生育进程的变化规律,以及氮素和株型对其的调控作用。随生育进程的推进,叶面积指数和群体茎蘖数的变化均呈单峰曲线,且分别于孕穗期和拔节期达到峰值。叶片净光合速率随生育进程先增加后下降,开花期达到峰值,于花后,叶片净光合速率逐渐下降。群体透光率的日变化呈抛物线状,且于14时达到最大值。不同株型小麦品种的叶面积指数、SPAD值、冠层绿叶面积和比叶重均存在差异,但群体茎蘖数差异不显著。于植株定形时,紧凑型品种宁麦9号和矮抗58、松散型品种淮麦17和中间型扬麦12的冠层垂直剖面绿叶面积的分布分别呈中部大上下两端小的直立“椭圆形”、上下基本一致的“柱形”、上小下大的“梯形”和上大下小的“倒梯形”。最大叶面积密度以紧凑型品种矮抗58的最大,中间型品种扬麦12的最小。施氮量对叶面积指数、SPAD值、比叶重均有显著影响。于孕穗期到成熟期,氮素对群体茎蘖数的调控效应显著。随施氮量增加,紧凑型品种宁麦9号和矮抗58的SPADmax、GDDmax、Rinf2和GDDinf2与中间型品种扬麦12和松散型品种淮麦17的变化趋势相反。施氮增加了小麦的茎蘖数、比叶重、净光合速率和最大叶面积密度,且提升了最大叶面积密度所在的叶层高度。
通过连续观测和定量分析不同处理下小麦干物质、氮素含量、籽粒产量及其构成,综合分析了不同株型小麦碳素、氮素积累与转运规律,以及籽粒产量构成差异。采用Richards和VP方程准确描述了不同处理下小麦干物质积累及其在不同器官间的分配比例动态。不同株型小麦干物质积累与分配对施氮量的响应差异主要表现在千物质积累的起始生长势、渐增和快增持续期,以及各器官的最大分配比例、分配比例最大下降速率和最大增长速率。不同器官的总糖转运量和转运率大小顺序分别为：茎鞘>叶片>穗部营养体,茎鞘>穗营养体>叶片。施氮量对4个株型小麦品种的叶、茎鞘和穗营养体可溶性总糖转运量的调控作用存在显著差异。不同株型小麦品种的茎鞘、穗营养体和叶片氮素输出量差异明显。施氮提高了4个株型小麦品种的花前氮积累量和成熟期氮积累量,但不同株型小麦品种在花后氮同化量上表现不一致。不同株型小麦产量差异主要来自千粒重和穗粒数,而穗数主要受氮素调节。
通过取样观测不同小麦品种和施氮水平下,株型形态特征、冠层结构特征、干物质和氮素积累与分配、及籽粒产量等指标,综合分析了籽粒产量及其构成因素与株型形态特征、冠层结构特征、碳氮积累与转运特征之间的相关关系。籽粒产量、千粒重与旗叶叶片特征呈显著正相关,但与倒二和倒三叶叶片特征的相关性较差；而穗粒数与顶部三张叶片特征均呈显著正相关。籽粒产量、千粒重和穗粒数与穗下节间长和倒四节间长均显著正相关,千粒重与倒二节间长、穗粒数与倒三节间长的相关性也达到显著水平。随生育进程的推进,籽粒产量和穗数与群体茎蘖数的相关性逐渐增强,而千粒重和穗粒数与群体茎蘖数的相关性未达到显著水平。籽粒产量、千粒重、穗粒数和穗数与花后干物质积累量、花前贮存物质转运量和花后物质积累对籽粒产量的贡献率(CPA)呈显著正相关,而与花前贮存物质对籽粒产量的贡献率(CTA)呈负相关。籽粒产量、千粒重和穗粒数与花前贮存物质转运率(TAR)呈显著正相关,而穗数与TAR呈显著负相关。籽粒产量、千粒重、穗数与氮转运量、氮转运率、花后氮同化对籽粒产量的贡献率呈显著正相关,而与花后氮转运对籽粒产量的贡献率呈负相关。在相同种植密度下,株型造成的产量差异是由于花后干物质同化量积累差异造成的千粒重和穗粒数差异。
The objective was to establish the regulation approaches of ideal plant type for high yielding. The split plot design was conducted with main plot of nitrogen fertilizer, and sub-plot of wheat cultivars differing in plant types (compact-high stalk type, Ningmai 9; compact-low stalk type, Aikang 58; Middle type, Yangmai 12; loosely type, Huaimai 17). This study focused on the effect of plant type and nitrogen fertilizer application rate on morpho-physiological characters of leaf and stem, canopy structure and light distribution, accumulation and translocation of carbon and nitrogen, grain yield, and the relationships among those characters.
The change patterns of leaf morphology with growth progress were characterized based on leaf morphological properties of main stems under different nitrogen rates with four wheat cultivars. The result showed that the trend of leaf length and width, and leaf angle looked like "S" shape with GDD. In contrast, Leaf SPAD increased gradually with GDD and reached the maximum at anthesis, and then decreased slowly. Leaf length and width, and leaf angle exhibited quadratic polynomial with different leaf position. Nevertheless, single leaf area had an "up-down-up-down" trend with leaf position. Leaf characteristics had a remarkable difference between cultivars at major growth stage. Nitrogen rate affected greatly leaf length and width, leaf angle, SPAD, and leaf area just at anthesis and grain filling period. Leaf length and width, leaf angle of different leaf position as affected by plant type and nitrogen rate had not the same response, but SPAD and distance between neighboring leaves of different leaf position had the same response. Difference of top two leaves leaf angle in Yangmai 12 and Huaimai 17 was larger than in Ningmai 12 and Aikang 58. The distance between neighboring leaves associated not only with plant type, but with also plant height. The distance between flag leaf and top 2nd leaf was the main difference of wheat cultivars with different plant type, as well as distance between top 2nd leaf and top 3rd leaf. Nitrogen rate strongly affected leaf length, leaf angle and SPAD, but had no effect on leaf width, leaf area, and distance between neighboring leaves at major growth stage. Nitrogen rate significantly affected leaf length of top three leaves and leaf width of top two leaves, but did not affect leaf width of top 3rn leaf. Nitrogen rate affected length and width of flag leaf more than that of top 2nd and 3rd leaf. Nitrogen rate increased leaf SPAD and leaf angle of top three leaves.
The pattern of stem characteristics and the effect of plant type and nitrogen rate on plant height, and length and width of internode was analyzed based on time-course observation on plant height and internode length and width of main stems. Plant height changed like "S" shape, and fitted well to Logistic equation during growing stage. Top four internode length was in the order: internode below ear> top 2nd internode> top 3rd internode> top 4th internode. The 4th internode width of Ningmai 9 and Aikang 58 and the top 2nd internode of Huaimai 17, and Yangmai 12 were the maximum. Nevertheless width of internode below ear was the minimum. Parameters derived from equation of plant height had great difference between nitrogen rates and plant types. Internode length and width correlated positively with nitrogen rate. Nitrogen rate strongly affected plant height at stem elongation period, full heading period and maturity, not yet at returning green. Nitrogen rate and plant type significantly affected internode length and width, but did not affect plant height index. Nitrogen rate and plant type influenced spike length, and nitrogen rate were positively related with spike length. So, there is a significant difference in plant height and internode of different plant type and nitrogen rate. Nitrogen rate greatly increase plant height, internode length and width, spike length.
Wheat canopy structure characteristics pattern was analyzed based on time-course observations on canopy characteristics under different nitrogen rates with four cultivars. The result showed that leaf area index and tiller number, which reached the maximum at booting stage and jointing stage, presented a single-peak curve, respectively. Net photosynthetic rate of leaf decreased gradually from flowering. Population light transmittance changed like parabola curves in diurnal course, and reached the maximum at noon. There was a great difference of plant type in LAI, SPAD, canopy green leaf area and specific leaf weight, not yet in population tiller number. The vertical distribution of green leaf area in four cultivars had significant difference. Nitrogen rate significantly affected LAI, SPAD and specific area weight. Nitrogen fertilization had a remarkable difference on population tiller number at booting and harvest. Ningmai 9 and Aikang 58 had a increase trend in SPADmax、GDDmax、Rinf2 and GDDinf2 with nitrogen rate increasing, but Yangmai 12 and Huaimai 17 had a reverse trend. Nitrogen rate enhanced tiller number, specific leaf weight, net photo synthetic rate and maximum leaf area density, and made level of maximum leaf area density gone up.
Carbon and nitrogen accumulation and translocation pattern and difference of grain yield in four cultivars were analyzed based on time-course observation on dry matter, nitrogen content, grain yield and its components. The result showed that relative dry matter and proportion of organs to biomass with relative thermal time from sowing fitted better to an Richards model and vapor pressure model, respectively. The dry matter accumulation duration of early stage decreased with nitrogen rate increasing under different wheat cultivars, whereas that of the third stage increased, and that of the second stage obviously differed from cultivars. Although the average growth rate Ra and maximum growth rate Rmax of dry matter increased with nitrogen rate increasing, that of dry matter dry mater decreased when nitrogen rate exceeded 150 kg ha-1, Transfer amount of soluble sugar ranked in the order of stem and sheath, leaf blades, and spike. Translocation rate of soluble sugar ranked in the order of stem and sheath, spike and leaf blades. Nitrogen fertilization affected remarkably transfer amount of soluble sugar in leaf, stem and sheath and spike of wheat cultivars. Nitrogen translocation amount of stem and sheath, spike and leaf in four wheat cultivars had significant differences. Nitrogen increased pre-anthesis nitrogen accumulation and nitrogen accumulation at maturity, but response of post-anthesis nitrogen assimilates in four wheat cultivars to nitrogen fertilization were different.
Dry matter and nitrogen accumulation and translocation, grain yield, correlation between grain yield and its components and plant type characteristics, canopy characters, carbon and nitrogen accumulation and translocation were analyzed based on time-course sampling observations. The result showed that grain yield and 1000-grain weight correlated significantly with the characteristics of flag leaf, but did not with that of the second and third leaf. Grain number per spike positively associated with characteristics of top three leaves. Moreover, grain yield,1000-grain weight, grain number per spike positively associated with length of upmost internode and second internode from top.1000-grain weight also significantly associated with length of second internode, and grain number per spike remarkably correlated with third internode. Grain yield and spike number closely associated with stem and tillers number, but 1000-grain and grain number per spike did not correlated with stem and tillers number. Grain yield,1000-grain, grain number per spike, and spike number significantly associated with post-anthesis dry matter accumulation, translocation amount of pre-anthesis stored assimilate, contribution of post-anthesis stored assimilate to grain yield, but had no relationship with contribution of pre-anthesis stored assimilate to grain yield. Grain yield,1000-grainweight and grain number per spike positively associated with translocation rate of pre-anthesis stored assimilate (TAR). Spike number with TAR showed a negative correlation. Grain yield,1000-grain and spike number with nitrogen translocation amount, nitrogen transportation efficiency, contribution of post-anthesis nitrogen assimilation to grain had a positive correlation, and with contribution of translocation amount of nitrogen accumulation to grain had a negative correlation.
本研究以不同氮素水平下、不同株型小麦品种的两年田间试验为基础,通过连续观测不同处理条件下小麦主茎叶片形态指标(包括叶长、叶宽、茎叶夹角、叶间距等),综合分析了小麦叶片形态指标随生育进程的变化规律,以及氮素和株型对叶片形态的调控效应。结果表明,随着生长度日的逐渐增加,小麦主茎叶片长度和宽度、茎叶夹角、单株叶面积均呈“S”形曲线变化,而叶片SPAD值,至开花期达最高值,随后逐渐下降。随着叶位的增加,小麦叶长和叶宽、茎叶夹角均符合二次多项式曲线,而单叶面积呈“升-降-升-降”的变化趋势。不同株型小麦品种顶部三张叶片的叶型指标差异主要表现在顶部三张叶片的叶长和SPAD值,旗叶与倒二叶间距及倒三叶基角。施氮量对不同株型小麦品种叶型指标的建成过程有显著影响,但对不同品种的影响程度有差异。
通过连续观测不同处理条件下小麦株高和主茎节间构成,综合分析了小麦株高及其构成、节间直径和长度随生育进程的变化规律,及氮素和株型对其的调控效应。结果表明,小麦株高随生长度日的动态伸长过程可以采用Logistic方程进行描述。小麦主茎顶四节的节间长大小顺序为：穗下节间>倒二节间>倒三节间>倒四节间,且均以穗下节间最细；紧凑型宁麦9号和矮抗58以倒四节间最粗,而松散型淮麦17和中间型扬麦12以倒二节间最粗。株高拟合方程的参数及衍生参数在不同施氮量和株型间存在显著差异。小麦主茎顶四节间的节间长和节间粗、穗长与施氮量呈正相关。在拔节、齐穗、灌浆和成熟期,施氮量对株高均有显著影响,而在返青期对株高的影响不显著。于齐穗、灌浆和成熟期,施氮量和株型对节间长和直径有显著影响,但对株高构成指数影响不显著。
基于不同氮素水平下、不同株型小麦品种叶面积指数、叶面积密度分布、透光率和光合速率等冠层特征的连续观测数据,综合分析了小麦冠层结构特征随生育进程的变化规律,以及氮素和株型对其的调控作用。随生育进程的推进,叶面积指数和群体茎蘖数的变化均呈单峰曲线,且分别于孕穗期和拔节期达到峰值。叶片净光合速率随生育进程先增加后下降,开花期达到峰值,于花后,叶片净光合速率逐渐下降。群体透光率的日变化呈抛物线状,且于14时达到最大值。不同株型小麦品种的叶面积指数、SPAD值、冠层绿叶面积和比叶重均存在差异,但群体茎蘖数差异不显著。于植株定形时,紧凑型品种宁麦9号和矮抗58、松散型品种淮麦17和中间型扬麦12的冠层垂直剖面绿叶面积的分布分别呈中部大上下两端小的直立“椭圆形”、上下基本一致的“柱形”、上小下大的“梯形”和上大下小的“倒梯形”。最大叶面积密度以紧凑型品种矮抗58的最大,中间型品种扬麦12的最小。施氮量对叶面积指数、SPAD值、比叶重均有显著影响。于孕穗期到成熟期,氮素对群体茎蘖数的调控效应显著。随施氮量增加,紧凑型品种宁麦9号和矮抗58的SPADmax、GDDmax、Rinf2和GDDinf2与中间型品种扬麦12和松散型品种淮麦17的变化趋势相反。施氮增加了小麦的茎蘖数、比叶重、净光合速率和最大叶面积密度,且提升了最大叶面积密度所在的叶层高度。
通过连续观测和定量分析不同处理下小麦干物质、氮素含量、籽粒产量及其构成,综合分析了不同株型小麦碳素、氮素积累与转运规律,以及籽粒产量构成差异。采用Richards和VP方程准确描述了不同处理下小麦干物质积累及其在不同器官间的分配比例动态。不同株型小麦干物质积累与分配对施氮量的响应差异主要表现在千物质积累的起始生长势、渐增和快增持续期,以及各器官的最大分配比例、分配比例最大下降速率和最大增长速率。不同器官的总糖转运量和转运率大小顺序分别为：茎鞘>叶片>穗部营养体,茎鞘>穗营养体>叶片。施氮量对4个株型小麦品种的叶、茎鞘和穗营养体可溶性总糖转运量的调控作用存在显著差异。不同株型小麦品种的茎鞘、穗营养体和叶片氮素输出量差异明显。施氮提高了4个株型小麦品种的花前氮积累量和成熟期氮积累量,但不同株型小麦品种在花后氮同化量上表现不一致。不同株型小麦产量差异主要来自千粒重和穗粒数,而穗数主要受氮素调节。
通过取样观测不同小麦品种和施氮水平下,株型形态特征、冠层结构特征、干物质和氮素积累与分配、及籽粒产量等指标,综合分析了籽粒产量及其构成因素与株型形态特征、冠层结构特征、碳氮积累与转运特征之间的相关关系。籽粒产量、千粒重与旗叶叶片特征呈显著正相关,但与倒二和倒三叶叶片特征的相关性较差；而穗粒数与顶部三张叶片特征均呈显著正相关。籽粒产量、千粒重和穗粒数与穗下节间长和倒四节间长均显著正相关,千粒重与倒二节间长、穗粒数与倒三节间长的相关性也达到显著水平。随生育进程的推进,籽粒产量和穗数与群体茎蘖数的相关性逐渐增强,而千粒重和穗粒数与群体茎蘖数的相关性未达到显著水平。籽粒产量、千粒重、穗粒数和穗数与花后干物质积累量、花前贮存物质转运量和花后物质积累对籽粒产量的贡献率(CPA)呈显著正相关,而与花前贮存物质对籽粒产量的贡献率(CTA)呈负相关。籽粒产量、千粒重和穗粒数与花前贮存物质转运率(TAR)呈显著正相关,而穗数与TAR呈显著负相关。籽粒产量、千粒重、穗数与氮转运量、氮转运率、花后氮同化对籽粒产量的贡献率呈显著正相关,而与花后氮转运对籽粒产量的贡献率呈负相关。在相同种植密度下,株型造成的产量差异是由于花后干物质同化量积累差异造成的千粒重和穗粒数差异。
The objective was to establish the regulation approaches of ideal plant type for high yielding. The split plot design was conducted with main plot of nitrogen fertilizer, and sub-plot of wheat cultivars differing in plant types (compact-high stalk type, Ningmai 9; compact-low stalk type, Aikang 58; Middle type, Yangmai 12; loosely type, Huaimai 17). This study focused on the effect of plant type and nitrogen fertilizer application rate on morpho-physiological characters of leaf and stem, canopy structure and light distribution, accumulation and translocation of carbon and nitrogen, grain yield, and the relationships among those characters.
The change patterns of leaf morphology with growth progress were characterized based on leaf morphological properties of main stems under different nitrogen rates with four wheat cultivars. The result showed that the trend of leaf length and width, and leaf angle looked like "S" shape with GDD. In contrast, Leaf SPAD increased gradually with GDD and reached the maximum at anthesis, and then decreased slowly. Leaf length and width, and leaf angle exhibited quadratic polynomial with different leaf position. Nevertheless, single leaf area had an "up-down-up-down" trend with leaf position. Leaf characteristics had a remarkable difference between cultivars at major growth stage. Nitrogen rate affected greatly leaf length and width, leaf angle, SPAD, and leaf area just at anthesis and grain filling period. Leaf length and width, leaf angle of different leaf position as affected by plant type and nitrogen rate had not the same response, but SPAD and distance between neighboring leaves of different leaf position had the same response. Difference of top two leaves leaf angle in Yangmai 12 and Huaimai 17 was larger than in Ningmai 12 and Aikang 58. The distance between neighboring leaves associated not only with plant type, but with also plant height. The distance between flag leaf and top 2nd leaf was the main difference of wheat cultivars with different plant type, as well as distance between top 2nd leaf and top 3rd leaf. Nitrogen rate strongly affected leaf length, leaf angle and SPAD, but had no effect on leaf width, leaf area, and distance between neighboring leaves at major growth stage. Nitrogen rate significantly affected leaf length of top three leaves and leaf width of top two leaves, but did not affect leaf width of top 3rn leaf. Nitrogen rate affected length and width of flag leaf more than that of top 2nd and 3rd leaf. Nitrogen rate increased leaf SPAD and leaf angle of top three leaves.
The pattern of stem characteristics and the effect of plant type and nitrogen rate on plant height, and length and width of internode was analyzed based on time-course observation on plant height and internode length and width of main stems. Plant height changed like "S" shape, and fitted well to Logistic equation during growing stage. Top four internode length was in the order: internode below ear> top 2nd internode> top 3rd internode> top 4th internode. The 4th internode width of Ningmai 9 and Aikang 58 and the top 2nd internode of Huaimai 17, and Yangmai 12 were the maximum. Nevertheless width of internode below ear was the minimum. Parameters derived from equation of plant height had great difference between nitrogen rates and plant types. Internode length and width correlated positively with nitrogen rate. Nitrogen rate strongly affected plant height at stem elongation period, full heading period and maturity, not yet at returning green. Nitrogen rate and plant type significantly affected internode length and width, but did not affect plant height index. Nitrogen rate and plant type influenced spike length, and nitrogen rate were positively related with spike length. So, there is a significant difference in plant height and internode of different plant type and nitrogen rate. Nitrogen rate greatly increase plant height, internode length and width, spike length.
Wheat canopy structure characteristics pattern was analyzed based on time-course observations on canopy characteristics under different nitrogen rates with four cultivars. The result showed that leaf area index and tiller number, which reached the maximum at booting stage and jointing stage, presented a single-peak curve, respectively. Net photosynthetic rate of leaf decreased gradually from flowering. Population light transmittance changed like parabola curves in diurnal course, and reached the maximum at noon. There was a great difference of plant type in LAI, SPAD, canopy green leaf area and specific leaf weight, not yet in population tiller number. The vertical distribution of green leaf area in four cultivars had significant difference. Nitrogen rate significantly affected LAI, SPAD and specific area weight. Nitrogen fertilization had a remarkable difference on population tiller number at booting and harvest. Ningmai 9 and Aikang 58 had a increase trend in SPADmax、GDDmax、Rinf2 and GDDinf2 with nitrogen rate increasing, but Yangmai 12 and Huaimai 17 had a reverse trend. Nitrogen rate enhanced tiller number, specific leaf weight, net photo synthetic rate and maximum leaf area density, and made level of maximum leaf area density gone up.
Carbon and nitrogen accumulation and translocation pattern and difference of grain yield in four cultivars were analyzed based on time-course observation on dry matter, nitrogen content, grain yield and its components. The result showed that relative dry matter and proportion of organs to biomass with relative thermal time from sowing fitted better to an Richards model and vapor pressure model, respectively. The dry matter accumulation duration of early stage decreased with nitrogen rate increasing under different wheat cultivars, whereas that of the third stage increased, and that of the second stage obviously differed from cultivars. Although the average growth rate Ra and maximum growth rate Rmax of dry matter increased with nitrogen rate increasing, that of dry matter dry mater decreased when nitrogen rate exceeded 150 kg ha-1, Transfer amount of soluble sugar ranked in the order of stem and sheath, leaf blades, and spike. Translocation rate of soluble sugar ranked in the order of stem and sheath, spike and leaf blades. Nitrogen fertilization affected remarkably transfer amount of soluble sugar in leaf, stem and sheath and spike of wheat cultivars. Nitrogen translocation amount of stem and sheath, spike and leaf in four wheat cultivars had significant differences. Nitrogen increased pre-anthesis nitrogen accumulation and nitrogen accumulation at maturity, but response of post-anthesis nitrogen assimilates in four wheat cultivars to nitrogen fertilization were different.
Dry matter and nitrogen accumulation and translocation, grain yield, correlation between grain yield and its components and plant type characteristics, canopy characters, carbon and nitrogen accumulation and translocation were analyzed based on time-course sampling observations. The result showed that grain yield and 1000-grain weight correlated significantly with the characteristics of flag leaf, but did not with that of the second and third leaf. Grain number per spike positively associated with characteristics of top three leaves. Moreover, grain yield,1000-grain weight, grain number per spike positively associated with length of upmost internode and second internode from top.1000-grain weight also significantly associated with length of second internode, and grain number per spike remarkably correlated with third internode. Grain yield and spike number closely associated with stem and tillers number, but 1000-grain and grain number per spike did not correlated with stem and tillers number. Grain yield,1000-grain, grain number per spike, and spike number significantly associated with post-anthesis dry matter accumulation, translocation amount of pre-anthesis stored assimilate, contribution of post-anthesis stored assimilate to grain yield, but had no relationship with contribution of pre-anthesis stored assimilate to grain yield. Grain yield,1000-grainweight and grain number per spike positively associated with translocation rate of pre-anthesis stored assimilate (TAR). Spike number with TAR showed a negative correlation. Grain yield,1000-grain and spike number with nitrogen translocation amount, nitrogen transportation efficiency, contribution of post-anthesis nitrogen assimilation to grain had a positive correlation, and with contribution of translocation amount of nitrogen accumulation to grain had a negative correlation.
引文
[1]Pimentel D, Harvey C, Resosudarmo P, Sinclair K, Kurz D, McNair M, Crist S, Shpritz L, Fitton L, Saffouri R, Blair R. Environmental and economic costs of soils erosion and conservation benefits[J]. Science,1995,267:1117-1123.
[2]程式华,曹立勇,庄杰云,吴伟明.关于超级稻品种培育的资源和基因利用问题[J].中国水稻科学,2009,23(3)：223-228.
[3]何中虎,张爱民.中国小麦育种研究进展[M].北京：中国科学技术出版社.2002
[4]魏燮中.我国古农书关于禾谷类作物株型及其生态表现的记载[J].农业考古,1987,1：292-299.
[5]Engledow F L, Wadham S M. Investigations on yield in the cereals[J]. The Journal of Agricultural Science,1923,13:390-439.
[6]陈友订,黄秋妹,张旭.水稻株型育种[M].上海：上海科学技术出版社.2005
[7]Donald C M. The breeding of crop ideotypes[J]. Euphytica,1968,17:385-403.
[8]Matsushima S. Theory and practice of rice cultivation[M]. Tokyo:Yokendo Publishing Co.1968
[9]Mick J J, Pearce R B. An ideotype of maize[J]. Euphytica,1975,24:613-623.
[10]杨守仁,陈温福,张龙步.水稻理想株型育种新动向[J].中国水稻科学,1988,2(3)：129-135.
[11]陈温福,徐正进,张龙步,杨守仁,韩显杰.水稻高产育种研究进展与前景[J].沈阳农业大学学报,1998,2：101-105.
[12]吴永成,周顺利,张永平,王志敏.节水高产小麦理想全株型探讨[J].干旱地区农业研究,2005,23(2)：126-129.
[13]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[14]徐庆章,李登海.玉米株型在高产育种中的作用[J].山东农业科学,1993,3：7-8.
[15]刘建栋,于强,吴乃元.黄淮海地区夏玉米株型结构特征分析[J].中国农业气象,2001,22(1)：6-14.
[16]张其鲁,陈香芝,张立全,李相奎,王家盛.小麦株型分类探讨[J].山东农业科学,2006,1：17-19.
[17]路明,周芳,谢传晓,李明顺,徐云碧,Warburton M,张世煌.玉米杂交种掖单13号的ssr连锁图谱构建与叶夹角和叶向值的qtl定位与分析[J].遗传,2007,29(9)：1131-1138.
[18]裴鑫德.玉米株型的判别松散型[J].华北农学报,1994,9(2)：1-8.
[19]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[20]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[21]郑丕尧.作物生理学导论[M].北京：北京农业大学出版社.1992
[22]Araus J L, Reynolds M P, Acevedo E. Leaf posture, grain yield, growth, leaf structure, and carbon isotope discrimination in wheat[J]. Crop Science,1993,33(6):1273-1279.
[23]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[24]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[25]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[26]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[27]范仲学,王璞,梁振兴,Zilkens M B, Claupein W.优化灌溉与施肥对冬小麦冠层结构的影响研究[J].中国生态农业学报,2005,13(3)：79-81.
[28]Singh P M,张国平,陈锦新.基因型和氮肥对冬小麦叶片形态建成和分蘖的影响[J].麦类作物学报,2000,20(1):28-23. (In English)
[29]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[30]李志勇,郭永新,王璞,翟志席,Zilkens M B.几种水氮模式处理下冬小麦冠层结构的差异[J].麦类作物学报,2007,27(6)：1085-1088.
[31]康祥波,钟阳和,张理,梅楠.冬小麦群体叶层结构的研究[J].河南职业技术师范学院学报,1990,18(3)：13-25.
[32]严威凯.关于小麦株形问题的观察与思考[J].作物杂志,1991,1：14-16.
[33]Annicchiarico P, Pecetti L. Developing a tall durum wheat plant type for semi-arid, mediterranean cereal-livestock farming systems[J]. Field Crops Research,2003,80:157-164.
[34]Marchetti M A. Potential impact of sheath blight on yield and milling quality of short-statured rice lines in the southern united states[J]. Plant Disease,1983,67:162-165.
[35]Sharma N R, Akanda S I, Shahjahan A K M. Development of sheath blight in short, tall, early and late maturing rice cultivars[J]. Bangladesh Journal of Botany,1995,24:143-146.
[36]Han Y P, Xing Y, Gu S, Chen Z, Pan X, Chen X. Effect of morphological traits on sheath blight resistance in rice[J]. Acta Botanica Sinica,2003,45:825-831.
[37]Savary S, Castilla N P, Elazegui F A, McLaren C G, Ynalvez M A, Teng P S. Direct and indirect effects of nitrogen supply and disease source structure on rice sheath blight spread[J]. Phytopathology,1995,85:959-1011.
[38]于经川,刘兆晔,姜鸿明,刘维正.小麦株高、穗幅宽和植株展开度的研究[J].莱阳农学院学报,2001,18(1)：7-11.
[39]张树榛,葛亚新.小麦理想株型研究：小麦株高梯度系列的株型、产量及品质性状的变化研究[J].北京农业大学学报,1990,16(2)：125-132.
[40]雷振生,林作楫.黄淮麦区冬小麦合理株型结构研究[J].华北农学报,1994,4：27-32.
[41]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[42]魏燮中,俞世蓉,宋荷仙,尚勋武.三个生态区小麦株型的比较分析[J].江苏农业学报,1991,7(1)：20-26.
[43]谢令琴,卢少源,吴同彦,王晓燕,常文锁,希慧军.冬小麦早代株高构成指数与产量关系的研究[J].河北农业大学学报,1996,19(1)：12-16.
[44]安呈峰,王延训,毕建杰,叶宝兴.高产小麦生育后期影响茎秆生长的生理因素与抗倒性的关系[J].山东农业科学,2008,7：1-4.
[45]赵万春,王红.小麦株高及其构成因素的遗传和相关性研究[J].麦类作物学报,2003,23(4)：28-31.
[46]吴玉娥,郜庆炉,杨文平,梁云娟,刘俊美.不同小麦品种冠层结构及产量研究[J].河南职业技术师范学院学报,2003,31(3)：1-4.
[47]朱新开,王祥菊,郭凯泉,郭文善,封超年,彭永欣.小麦倒伏的茎秆特征及对产量与品质的影响[J].麦类作物学报,2006,26(1)：87-92.
[48]吴同彦,谢令琴,杨学举,张彩英,陈荣芬.小麦株高构成因素与产量及其他性状相关性的研究[J].河北农业大学学报,2002,25(3)：10-12.
[49]Sharma R C. Tiller mortality and its relationship to grain yield in spring wheat[J]. Field Crops Research,1995,41:55-60.
[50]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var. Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103:25-35.
[51]Jones H G, Kirby E J M. Effects of manipulation of number of tillers and water supply on grain yield in barley[J]. The Journal of Agricultural Science,1977,88:391-397.
[52]Islam T M T, Sedgley R H. Evidence for a "uniculm effect" in spring wheat (triticum aestivum l.) in a mediterranean environment[J]. Euphytica,1981,30:277-282.
[53]李少昆,王崇桃.作物株型和冠层结构信息获取与表达的方法[J].石河子大学学报(自然科学版),1997,1(3)：250-256.
[54]张艳敏,李晋生,钱维朴,黄德明.小麦冠层结构与光分布研究[J].华北农学报,1996,11(1)：54-58.
[55]张立全,张其鲁.小麦株型与田间透光率关系研究[J].潍坊学院学报,2006,6(6)：95-98.
[56]王之杰,郭天财,朱云集,王纪华,赵明.超高产小麦冠层光辐射特征的研究[J].西北植物学报,2003,23(10)：1657-1662.
[57]孙旭生,林琪,李玲燕,姜雯,翟延举.施氮量对超高产小麦生育后期光合特性及产量的影响[J].植物营养与肥料学报,2008,14(5)：840-844.
[58]赵春江,吴华瑞,王纪华,黄文江.田间小麦叶面积空间分布数学模型的建立与应用[J].中国农业科学,2004,37(2)：196-200.
[59]李萌,隋娜,王鹏,赵世杰,孟庆伟,田纪春.生育后期冠层结构对超高产小麦品种(系)光能利用及产量的影响[J].麦类作物学报,2009,29(2)：279-282.
[60]司纪升,王法宏,李升东,冯波,孙令安.不同种植方式对小麦群体质量和产量结构的影响[J].麦类作物学报,2006,26(6)：136-139.
[61]李春明,熊淑萍,赵巧梅,杨颖颖,马新明.有机无机肥配施对小麦冠层结构、产量和蛋白质含量的影响[J].中国农业科学,2008,41(12)：4287-4293.
[62]杨文平,郭天财,刘胜波,王晨阳,王永华,马冬云.行距配置对‘兰考矮早八’小麦后期群体冠层结构及其微环境的影响[J].植物生态学报,2008,32(2)：485-490.
[63]Rasmusson D C. A plant breeder's experience with ideotype breeding[J]. Field Crops Research, 1991,26:191-200.
[64]严威凯.关于小麦株型的观察与思考[J].作物杂志,1991,(1)：14-16.
[65]Yin X, Kropff M J, Goudriaan J, Stam P. A model analysis of yield differences among recombinant inbred lines in barley[J]. Agronomy Journal,2000,92:114-120.
[66]Baker C J, Berry P M, Spink J H, Sylvester-Bradley R, Griffin J M, Scott R K, Clare R W. A method for the assessment of the risk of wheat lodging [J]. Journal of Theoretical Biology,1998, 194(4):587-603.
[67]Berry P M, Sterling M, Baker C J, Spink J H, Sparkes D L. A calibrated model of wheat lodging compared with field measurements[J]. Agricultural and Forest Meteorology,2003,19(4):445-451.
[68]李韬,徐辰武,丁永辉,胡治球,孙长森,顾世梁.一些小麦品种主要株型指标的初步分析[J].扬州大学学报：农业与生命科学版,2003,34(2)：40-43,48.
[69]张其鲁.潍麦8号的株型特征及小麦理想株型研究[D].北京：中国农业大学.2004
[70]吴同彦,谢令琴,葛淑俊,张彩英,王静华,王宝山.小麦营养器官与产量性状相关性的研究[J].河北农业大学学报,2001,24(4)：11-13.
[71]Kropff M J. Application of systems approaches at the field levels[M]. Kluwer Academic Publisher. 1996
[72]Xu Z Z, Yu Z W, Wang D, Zhang Y L. Nitrogen accumulation and translocation for winter wheat under different irrigation regimes[J]. Agronomy and Crop Science,2005,191:439-449.
[73]Ledent J F, Moss D N. Relation of morphological characters and shoot yield in wheat[J]. Crop Science,1979,19(4):445-451.
[74]Yunusa I A M, Siddique K H M, Belford R K, Karimi M M. Effect of canopy structure on efficiency of radiation interception and use in spring wheat cultivars during the pre-anthesis period in a mediterranean-type environment[J]. Field Crops Research,1993,35(2):113-122.
[75]Latiri-Souki K, Nortcliff S, Lawlor D W. Nitrogen fertilizer can increase dry matter, grain production and radiation and water use efficiencies for durum wheat under semi-arid conditions[J]. European Journal of Agronomy,1998,9(1):21-34.
[76]张定一,党建友,王姣爱,裴雪霞,杨武德,苗果园.施氮量对不同品质类型小麦产量、品质和旗叶光合作用的调节效应[J].植物营养与肥料学报,2007,13(4)：535-542.
[77]王月福,姜东,于振文.氮素水平对小麦籽粒产量和蛋白质含量的影响及其生理基础[J].中国农业科学,2003,36(5)：513-520.
[78]周凌云,李卫民.氮素营养对干旱地区小麦光合生理的影响[J].土壤通报,2003,34(3)：195-197.
[79]陆增根,戴廷波,姜东,荆奇,吴正贵,周培南,曹卫星.氮肥运筹对弱筋小麦群体指标与产量和品质形成的影响[J].作物学报,2007,33(4)：590-597.
[80]王月福,于振文,李尚霞,余松烈.土壤肥力和施氮量对小麦根系氮同化及子粒蛋白质含量的影响[J].植物营养与肥料学报,2003,9(1)：39-44.
[81]Pinthus M J. Lodging in wheat, barley, and oats:The phenomenon, its causes, and preventive measures. In Advances in agronomy, vol. Volume 25:Academic Press.209-263.1974.
[82]Kheiralla K A, Mehdi E E, Dawood R A. Evaluation of some wheat cultivars for traits related to lodging resistance under different levels of nitrogen[J]. Assiut Journal of Agricultural Science,1993, 24:257-271.
[83]Crook M J, Ennos A R. The effect of nitrogen and growth regulators on stem and root characteristics associated with lodging in two cultivars of winter wheat[J]. Journal of Experimental Botany,1995,46(8):931-938.
[84]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Growth and morphology of spring wheat (triticum aestivum 1.) culms and their association with lodging: Effects of genotypes, n levels and ethephon[J]. Field Crops Research,2003,84(3):271-290.
[85]张军,戴其根,张洪程,许轲,霍中洋.追氮时期对中筋小麦群体质量及品质的调节效应[J].江苏农业科学,2003,5：34-36.
[86]王月福,于振文,李尚霞,张永丽.不同土壤肥力下强筋小麦适宜施氮量的研究[J].山东农业科学,2001,5：14-15.
[87]Aude B, Christophe L, Christine B, Marie-Helene J. Nitrogen remobilization during grain filling in wheat: Genotypic and environmental effects[J]. Crop Science,2005,45(3):1141.
[88]Arduini I, Masoni A, Ercoli L, Mariotti M. Grain yield, and dry matter and nitrogen accumulation and remobilization in durum wheat as affected by variety and seeding rate[J]. European Journal of Agronomy,2006,25(4):309-318.
[89]Muurinen S, Kleemola J, Peltonen-Sainio P. Accumulation and translocation of nitrogen in spring cereal cultivars differing in nitrogen use efficiency [J]. Agronomy Journal,2007,99(2):441-449.
[90]Ercoli L, Lulli L, Mariotti M, Masoni A, Arduini I. Post-anthesis dry matter and nitrogen dynamics in durum wheat as affected by nitrogen supply and soil water availability[J]. European Journal of Agronomy,2008,28(2):138-147.
[91]王月福,于振文,李尚霞,余松烈.氮素营养水平对小麦开花后碳素同化、运转和产量的影响[J].麦类作物学报,2002,22(2)：55-59.
[92]王月福,于振文,李尚霞,余松烈.土壤肥力和施氮量对小麦氮素吸收运转及籽粒产量和蛋白质含量的影响[J].应用生态学报,2003,14(11)：1868-1872.
[93]沈建辉,戴廷波,荆奇,姜东,曹卫星,郭文善,王龙俊.施氮时期对专用小麦干物质和氮素积累、运转及产量和蛋白质含量的影响[J].作物学报,2004,24(1)：55-58.
[94]吴清丽,高茂盛,廖允成,温晓霞.氮素对冬小麦光合物质贮运及籽粒灌浆进程的影响[J].干旱地区农业研究,2009,27(2)：120-124.
[95]武际,郭熙盛,王允青,汪建来,杨晓虎.氮钾配施对弱筋小麦氮、钾养分吸收利用及产量和品质的影响[J].植物营养与肥料学报,2007,13(6)：1054-1061.
[96]杨延兵,高荣岐,尹燕枰,吴承来,李圣福.不同品质小麦氮素分配及利用率的15n示踪研究[J].麦类作物学报,2008,28(5)：830-835.
[97]同延安,赵营,赵护兵,樊红柱.施氮量对冬小麦氮素吸收、转运及产量的影响[J].植物营养与肥料学报,2007,13(1)：64-69.
[98]石玉,于振文,王东,李延奇,王雪.施氮量和底追比例对小麦氮素吸收转运及产量的影响[J].作物学报,2006,32(12)：1860-1866.
[99]于振文,潘庆民,姜东,张永丽,王东.9000 kg/公顷小麦施氮量与生理特性分析[J].作物学报,2003,29(1)：37-43.
[100]王小燕,于振文.不同施氮量条件下灌溉量对小麦氮素吸收转运和分配的影响[J].中国农业科学,2008,41(10)：3015-3024.
[101]吴海卿,段爱旺,杨传福.冬小麦对不同土壤水分的生理和形态响应[J].华北农学报,2000,5(1)：92-96.
[102]冯广龙,罗远培,杨培岭.节水灌溉对冬小麦干物质分配、灌浆及水分利用率的影响[J].华北农学报,1998,13(2)：11-17.
[103]胡梦芸,张正斌,徐萍,董宝娣,李魏强,李景娟.亏缺灌溉下小麦水分利用效应与光合产 物积累运转的相关研究[J].作物学报,2007,33(10)：1711-1719.
[104]Plaut Z, Butow B J, Blumenthal C S, Wrigley C W. Transport of dry matter into developing wheat kernels and its contribution to grain yield under post-anthesis water deficit and elevated temperature[J]. Field Crops Research,2004,86(2-3):185-198.
[105]王立秋,曹敬山,靳占忠.春小麦产量及品质的水肥效应研究[J].干旱地区农业研究,1997,15(1)：58-63.
[106]王晓燕,于振文.不同施氮量条件下灌溉量对小麦氮素吸收转运和分析的影响[J].中国农业科学,2008,41(10)：3105-3024.
[107]刘萍,郭文善,徐月明,封超年,朱新开,彭永欣.种植密度对中、弱筋小麦籽粒产量和品质的影响[J].麦类作物学报,2006,26(5)：117-121.
[108]吴兰云,周得宝,徐茂林.淮北地区高产小麦的氮肥和密度效应研究[J].中国农学通报,2008,24(7)：153-157.
[109]郭天财,盛坤,冯伟,徐丽娜,王晨阳.种植密度对两种穗型小麦品种分蘖期茎蘖生理特性的影响[J].西北植物学报,2009,29(2)：350-355.
[110]屈会娟,李金才,沈学善,魏凤珍,王成雨,郅胜军.种植密度和播期对冬小麦品种兰考矮早八干物质和氮素积累与转运的影响[J].作物学报,2009,35(1)：124-131.
[111]武兰芳,欧阳竹.不再种植密度下两种穗型小麦叶片光合特性的变化[J].麦类作物学报,2008,28(4)：618-625.
[112]刘万代,杜沛鑫,尹钧,陈现勇.种植密度对超高产小麦豫麦49-198籽粒产量及品质的影响[J].安徽农业科学,2008,36(35)：15402-15404.
[113]于振文,岳寿松,沈成国,余松烈,Egli D B, Sanford V V.不同密度对冬小麦开花后叶片衰老和粒重的影响[J].作物学报,1995,21(4)：412-418.
[114]徐月明.中弱筋小麦优质高产群体质量和株型指标与生理基础研究[D].扬州大学.2004
[115]郭天财,查菲娜,马冬云,宋晓,岳艳军.种植密度对两种穗型冬小麦品种干物质和氮素积累、运转及产量的影响[J].华北农学报,2007,22(6)：152-156.
[116]张晓龙.小麦品种籽粒灌浆研究[J].作物学报,1982,8(2)：87-93.
[117]李金才.品种和播种密度灌浆特性及产量影响的研究[J].安徽农业大学学报,1996,23(4)：461-465.
[118]杨永光,张维成.播量对小麦产量和籽粒营养品质的影响[J].河南职业技术师范学院学报,1989,17(3)：113-116.
[119]董爱民,甘淼,王秀玉,冯新忠,韩林,陈建峰.不同肥力与密度条件下对小麦群体结构及产量的研究[J].河南职业技术师范学院学报,2004,32(2)：12-14.
[1]魏燮中.小麦株型结构分析与产量育种咨询系统[M].南京：东南大学出版社.1991
[2]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[3]张其鲁.潍麦8号的株型特征及小麦理想株型研究[D].北京：中国农业大学.2004
[4]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[5]王士红,荆奇,戴廷波,姜东,曹卫星.不同年代冬小麦品种旗叶光合特性和产量的演变特征[J].应用生态学报,2006,19(6)：1255-1260.
[6]李合生.植物生理生化实验原理和技术[M].北京：高等教育出版社.2000
[7]Gallagher J N. Field studies of cereal leaf growth: Ⅰ. initiation and expansion in relation to temperature and ontogeny[J]. Journal of Experimental Botany,1979,30(4):625-636.
[8]Draper N R, Smith H. Applied regression analysis[M]. John Wiley and Sons.1998
[9]SAS Institute. SAS/STAT 9.1 User's Guide[M]. Cary, NC.2004
[1]任书杰,李世清,王全九,李生秀.栽培模式、施氮和品种对冬小麦冠层结构和产量的影响[J].生态学杂志,2006,25(12)：1449-1454.
[2]Donald C M. The breeding of crop ideotypes[J]. Euphytica,1968,17:385-403.
[3]严威凯.关于小麦株形问题的观察与思考[J].作物杂志,1991,1：14-16.
[4]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[5]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[6]Araus J L, Reynolds M P, Acevedo E. Leaf posture, grain yield, growth, leaf structure, and carbon isotope discrimination in wheat[J]. Crop Science,1993,33(6):1273-1279.
[7]林忠辉,周允华,王辉民,张宜光.青藏高原冬小麦冠层几何结构、光截获及其光合潜能的影响[J].生态学报,1998,18(4)：392-398.
[8]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[9]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[10]Briggs K G, Aytenfisu A. Relationships between morphological characters above the flag leaf node and grain yield in spring wheats[J]. Crop Science,1980,20(3):350-354.
[11]Ledent J F, Moss D M. Relation of morphological characters and shoot yield in wheat[J]. Crop Science,1979,19(4):445-451.
[12]Ishag H M, Mohamed B A, Ishag K H M. Leaf development of spring wheat cultivars in an irrigated heat-stressed environment J]. Field Crops Research,1998,58(3):167-175.
[13]Monneveux P, Reynolds M P, Gonzaez-Santoyo H, Pena R J, Mayr L, Zapata F. Relationships between grain yield, flag leaf morphology, carbon isotope discrimination and ash content in irrigated wheat[J]. Journal of Agronomy and Crop Science,2004,190(6):395-401.
[14]Blake N K, Lanning S P, Martin J M, Sherman J D, Talbert L E. Relationship of flag leaf characteristics to economically important traits in two spring wheat crosses[J]. Crop Science,2007, 47(2):491-494.
[15]王法宏,杨洪宾,徐成忠,李升东,司纪升.垄作栽培对小麦植株形态和产量性状的影响[J].作物学报,2007,33(6)：1038-1040.
[16]Schabenberger O, Pierce F J. Contemporary statistical models for the plant and soil sciences [M]. New York: CRC Press.2002
[17]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[18]陈国庆.小麦形态建成模拟及可视化研究[D].山东农业大学.2004
[19]常丽英.水稻植株形态建成的模拟模型研究[D].南京农业大学.2007
[20]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[21]王义芹,杨兴洪,李滨,童依平,李振声.小麦叶面积及光合速率与产量关系的研究[J].华北农学报,2008,23(增刊)：10-15.
[22]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[23]Singh P M, Zhang G P, Chen J X. Influence of genotypes and nitorgen fertilization on leaf morphogenesis and tillering behaviors in winter wheat[J]. Journal of Triticeae Crops,2000,20(1): 28-23.
[24]李志勇,郭永新,王璞,翟志席,Zilkens M B.几种水氮模式处理下冬小麦冠层结构的差异[J].麦类作物学报,2007,27(6)：1085-1088.
[25]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[26]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[27]伍维模,李世清,王瑞军,邵明安.两种施氮水平下不同基因型冬小麦叶片光合特性与形态差异的研究[J].中国生态农业学报,2006,14(4)：76-81.
[28]Ledent J F. Relationships between grain yield and morphological characters in the winter wheat genotypes of the belgian national list[J]. Biologia Plantarum,1979,21(3):161-169.
[29]Hansen K A, Martin J M, Lanning S P, Talbert L E. Correlation of genotype performance for agronomic and physiological traits in space-planted versus densely seeded conditions[J]. Crop Sci, 2005,45(3):1023-1028.
[30]佟汉文,黄荣华,刘易科,朱展望,张宇庆,秦金成,付汉红,高春保.小麦新品种农艺性状与产量的相关及通径分析[J].湖北农业科学,2008,2008(47)：7.
[31]余泽高,王孝刚.小麦茎叶性状与穗部性状相关性的探讨[J].湖北农学院学报,1998,(1)：8-11.
[32]贾继增.小麦粒重与植株性状相关因素的统计分析[J].作物学报,1984,10(3)：201-205.
[1]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[2]欧阳由男,李春生,章善庆,王会民,朱练峰,禹盛苗,金千瑜,张国平.光周期和有效积温对水稻分蘖角度动态变化的影响[J].应用生态学报,2009,20(5)：1099-1104.
[3]闵东红,翟耀锋,王辉,孟超敏,崔耀锋,李学军,曹宁,于新智.不同株高小麦品种抗倒伏性与其亚性状及产量相关性研究[J].麦类作物学报,2001,21(4)：76-79.
[4]武仙山,王正航,昌小平,景蕊莲.用株高旱胁迫系数分析小麦发育中的抗旱性动态[J].作物学报,2008,34(11)：2010-2018.
[5]赵万春,王红.小麦株高及其构成因素的遗传和相关性研究[J].麦类作物学报,2003,23(4)：28-31.
[6]于经川,刘兆晔,姜鸿明,刘维正,马淑丽.小麦株高整齐度的初步研究[J].莱阳农学院学报,2002,19(1)：20-22.
[7]吴同彦,谢令琴,杨学举,张彩英,陈荣芬.小麦株高构成因素与产量及其他性状相关性的研究[J].河北农业大学学报,2002,25(3)：10-12.
[8]于经川,刘兆晔,姜鸿明,刘维正.小麦株高、穗幅宽和植株展开度的研究[J].莱阳农学院学报,2001,18(1)：7-11.
[9]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var. Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103:25-35.
[10]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Growth and morphology of spring wheat(triticum aestivum 1.) culms and their association with lodging: Effects of genotypes, n levels and ethephon[J]. Field Crops Research,2003,84(3):271-290.
[11]Sharma R C. Tiller mortality and its relationship to grain yield in spring wheat[J]. Field Crops Research,1995,41(1):55-60.
[12]Weiss A, Baenziger P S, McMaster G S, Wilhelm W W, Al Ajlouni Z I. Quantifying phenotypic plasticity using genetic information for simulating plant height in winter wheat[J]. NJAS Wageningen Journal of Life Sciences,2009,57(1):59-64.
[13]白云凤,李希陵.小麦几个数量性状不同生长时期的基因效应分析[J].华北农学报,1997,12(3)：11-16.
[14]蒋向辉,佘朝文,刘伟,何雪梅.青蒿株高生长与分枝特性的曲线方程及其拟合性研究[J].作物杂志,2008,3：46-49.
[15]张殿顺,董翔云,刘树庆.不同施氮水平对春小麦生长发育及其氮素代谢指标的影响[J].华北农学报,2006,21(增刊2)：42-45.
[16]Slafer G A, Halloran G M, Connor D J. Influence of photoperiod on culm length in wheat[J]. Field Crops Research,1995,40(2):95-99.
[17]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Effect of planting methods and n rates on lodging, morphological characters of culm and yield in spring wheat varieties[J]. Cereal Research Communications,2002,30(3-4):431-438.
[18]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[19]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[20]张娟.超高产小麦品种冠层结构对产量的影响及其遗传研究[D].河南农业大学.2000
[21]胡延吉,兰进好.不同时期小麦主栽品种冠层结构研究[J].中国农业气象,1999,20(1)：11-14.
[22]王淑俭.小麦株高与产量性状相关性的初步研究[J].河南农林科技,1982,7：1-3.
[23]张耀中.小麦株高与产量性状的相关分析[J].河南农业技术师范学院学报,1988,2(1)：59-60.
[24]贾继增.小麦粒重与植株性状相关因素的统计分析[J].作物学报,1984,10(3)：201-205.
[25]庄萍萍,李伟,魏育明,颜泽洪,郑有良.波斯小麦农艺性状相关性及主成分分析[J].麦类作物学报,2006,26(4)：11-14.
[1]胡延吉,兰进好.山东省冬小麦品种冠层结构及光截获的研究[J].中国农业气象,2001,22(3)：28-32.
[2]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[3]吕丽华,赵明,赵久然,陶洪斌,王璞.不同施氮量下夏玉米冠层结构及光合特性的变化不同施氮量下夏玉米冠层结构及光合特性的变化[J].中国农业科学,2008,41(9)：2624-2632.
[4]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[5]胡延吉,兰进好,赵坦方,高法振.不同穗型的两个冬小麦品种冠层结构及光合特性的研究[J].作物学报,2000,26(6)：905-912.
[6]赵会杰,李有,邹琦.两个不同穗型小麦品种的冠层辐射和光合特征的比较研究[J].作物学报,2002,28(5)：654-659.
[7]任书杰,李世清,王全九,李生秀.栽培模式、施氮和品种对冬小麦冠层结构和产量的影响[J].生态学杂志,2006,25(12)：1449-1454.
[8]杨文平,郭天财,刘胜波,王晨阳,王永华,马冬云.行距配置对‘兰考矮早八’小麦后期群体冠层结构及其微环境的影响[J].植物生态学报,2008,32(2)：485-490.
[9]熊淑萍,杨颖颖,李春明,马新明,姬兴杰,徐光辉.不同肥料处理对豫麦34冠层结构特征及产量的影响[J].麦类作物学报,2008,28(2)：276-280.
[10]李春明,熊淑萍,赵巧梅,杨颖颖,马新明.有机无机肥配施对小麦冠层结构、产量和蛋白质含量的影响[J].中国农业科学,2008,41(12)：4287-4293.
[11]吴玉娥,郜庆炉,薛香,梁云娟,韩占江.行距对超高产小麦冠层结构及产量构成的影响[J].河南农业科学,2005,9：16-20.
[12]范仲学,王璞,梁振兴,Zilkens M B, Claupein W.优化灌溉与施肥对冬小麦冠层结构的影响研究[J].中国生态农业学报,2005,13(3)：79-81.
[13]张殿顺,董翔云,刘树庆.不同施氮水平对春小麦生长发育及其氮素代谢指标的影响[J].华北农学报,2006,21：42-45.
[14]陆增根,戴廷波,姜东,荆奇,吴正贵,周培南,曹卫星.氮肥运筹对弱筋小麦群体指标与产量和品质形成的影响[J].作物学报,2007,33(4)：590-597.
[15]蔡瑞国,张敏,戴忠民,田雷,王振林.施氮水平对优质小麦旗叶光合特性和子粒生长发育的影响[J].植物营养与肥料学报,2006,12(1)：49-55.
[16]周春菊,张嵩午,王林权,苗芳.施肥对小麦冠层温度的影响及其与生物学性状的关联[J].生 态学报,2005,25(1)：18-22.
[17]孙旭生,林琪,赵长星,翟延举,吴钢,Jaleel C A.施氮量对超高产冬小麦灌浆期旗叶光响应曲线的影响[J].生态学报,2009,29(3)：1428-1437.
[18]赵长星,马东辉,王月福,林琪.施氮量和花后土壤含水量对小麦旗叶衰老及粒重的影响[J].应用生态学报,2008,19(11)：2388-2393.
[19]Hosoi F, Omasa K. Estimating vertical plant area density profile and growth parameters of a wheat canopy at different growth stages using three-dimensional portable lidar imaging[J].ISPRS Journal of Photogrammetry and Remote Sensing,2009,64(2):151-158.
[20]严威凯,王新中.小麦冠层结构中叶面积的层次构成与分析[J].陕西农业科学,1991,3：36-37.
[21]张艳敏,李晋生,钱维朴,黄德明.小麦冠层结构与光分布研究[J].华北农学报,1996,11(1)：54-58.
[22]张娟.超高产小麦品种冠层结构对产量的影响及其遗传研究[D].河南农业大学.2000
[23]Shi Z L, Yin M, Jing Q, Jiang D, Cao W X, Dai T B. Vertical distribution of canopy nitrogen and its relationship with grain protein in winter[J]. Journal of Triticeae Crops,2009,29(2):289-293.
[24]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[25]杨长明,刘敏华,丁超尘,杨林章.三个水稻品种的冠层结构比较研究[J].安徽农业大学学报,2001,28(3)：276-280.
[26]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[27]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[28]王谦,陈景玲,孙治强.LAI-2000冠层分析仪在不同植物群体光分布特征研究中的应用[J].中国农业科学,2006,39(5)：922-927.
[29]章家恩.作物群体建构的生态环境效应及其优化探讨[J].生态科学,2000,19(1)：30-35.
[30]Lotscher M, Stroh K, Schnyder H. Vertical leaf nitrogen distribution in relation to nitrogen status in grassland plants[J]. Annals of Botany,2003,92(5):679-688.
[31]Bertheloot J, Martre P, Andrieu B. Dynamics of light and nitrogen distribution during grain filling within wheat canopy[J]. Plant Physiology,2008,148(3):1707-1720.
[32]Dreccer M F, van Oijen M, Schapendonk A H C M, Pot C S, Rabbinge R. Dynamics of vertical leaf nitrogen distribution in a vegetative wheat canopy: Impact on canopy photosynthesis[J]. Annals of Botany,2000,86(4):821-831.
[33]Kull O. Acclimation of photosynthesis in canopies:Models and limitations[J]. Oecologia,2002, 133(3):267-279.
[34]Pommel B, Gallais A, Coque M, Quiller I, Hirel B, Prioul J L, Andrieu B, Floriot M. Carbon and nitrogen allocation and grain filling in three maize hybrids differing in leaf senescence[J]. European Journal of Agronomy,2006,24(3):203-211.
[35]Field C. Allocating leaf nitrogen for the maximization of carbon gain: Leaf age as a control on the allocation program[J]. Oecologia,1983,56(2):341-347.
[36]Hirose T, Werger M J A. Maximizing daily canopy photosynthesis with respect to the leaf nitrogen allocation pattern in the canopy[J]. Oecologia,1987,72(4):520-526.
[37]王之杰,王纪华,黄文江,马智红,赵明.冬小麦冠层不同叶层和茎鞘氮素与籽粒品质关系的研究[J].中国农业科学,2003,36(12)：1462-1468.
[38]王之杰,王纪华,黄文江,马镥宏,王北洪,赵春江,赵明.冬小麦叶片氮素时空分布特征及其与籽粒品质的关系[J].作物学报,2004,30(7)：700-707.
[39]石祖梁,殷美,荆奇,姜东,曹卫星,戴廷波.冬小麦冠层氮素垂直分布特征及其与籽粒蛋白质的关系[J].麦类作物学报,2009,29(2)：289-293.
[40]秦晓东,戴廷波,荆奇,姜东,曹卫星.冬小麦叶片氮含量时空分布及其与植株氮营养状况的关系[J].作物学报,2006,31(11)：1717-1722.
[41]Milroy S P, Bange M P, Sadras V O. Profiles of leaf nitrogen and light in reproductive canopies of cotton (gossypium hirsutum)[J]. Annals of Botany,2001,87(3):325-333.
[42]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[43]Lv L H, Zhao M, Zhao J R, Tao H B, P W. Canopy structure and photosynthesis of summer maize under different nitrogen fertilizer application rates[J].中国农业科学,2008,41(9):2624-2632.
[1]杨文平.行距和密度对冬小麦冠层结构、微环境及碳氮代谢的影响[D].河南农业大学.2008
[2]姜东,于振文,等.高产冬小麦茎中果聚糖代谢及氮素水平的调控[J].作物学报,2002,28(1)：79-85.
[3]韩占江,郜庆炉,薛香,吴玉娥,敖玲.不同小麦品种干物质积累、分配与产量的关系[J].贵州农业科学,2008,36(6)：14-16.
[4]张利,隋新霞,王羽,樊庆琦,李根英,楚秀生,黄承彦.不同熟性小麦品种的干物质积累和分配规律[J].山东农业科学,2007,6：54-56.
[5]吴清丽,高茂盛,廖允成,温晓霞.氮素对冬小麦光合物质贮运及籽粒灌浆进程的影响[J].干旱地区农业研究,2009,27(2)：120-124.
[6]彭正萍,王艳群,刘淑桥,王红,王蕾,薛世川.不同施肥处理对冬小麦干物质积累及土壤养分垂直分布的影响[J].中国农业科技导报,2007,9(6)：95-99.
[7]马东辉,王月福,周华,孙虎.氮肥和花后土壤含水量对小麦干物质积累、运转及产量的影响[J].麦类作物学报,2007,27(5)：847-851.
[8]屈会娟,李金才,沈学善,魏凤珍,王成雨,郅胜军.种植密度和播期对冬小麦品种兰考矮早八干物质和氮素积累与转运的影响[J].作物学报,2009,35(1)：124-131.
[9]伍维模,李世清,王瑞军,邵明安.两种施氮水平下不同基因型冬小麦叶片光合特性与形态差异的研究[J].中国生态农业学报,2006,14(4)：76-81.
[10]武际,郭熙盛,王允青,汪建来,杨晓虎.氮钾配施对弱筋小麦氮、钾养分吸收利用及产量和品质的影响[J].植物营养与肥料学报,2007,13(6)：1054-1061.
[11]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[12]Schabenberger O, Pierce F J. Contemporary statistical models for the plant and soil sciences[M]. Boca Raton:Taylor & Francis/CRC Press.2001
[13]朱庆森,曹显祖,骆亦其.水稻籽粒灌浆的生长分析[J].作物学报,1988,14(3)：182-193.
[14]顾世梁,朱庆森,杨建昌,彭少兵.不同水稻材料籽粒灌浆特性的分析[J].作物学报,2001,27(1)：7-14.
[15]Darroch B A, Baker R J. Grain filling in three spring wheat genotypes:Statistical analysis[J]. Crop Scinece,1990,30:525-529.
[16]沈建辉,戴廷波,荆奇,姜东,曹卫星,郭文善,王龙俊.施氮时期对专用小麦干物质和氮素积累、运转及产量和蛋白质含量的影响[J].麦类作物学报,2004,24(1)：55-58.
[17]王月福,于振文,李尚霞,余松烈.氮素营养水平对小麦开花后碳素同化、运转和产量的影响[J].麦类作物学报,2002,22(2)：55-59.
[18]姚素梅,康跃虎,刘海军.喷灌与地面灌溉冬小麦干物质积累、分配和运转的比较研究[J].干旱地区农业研究,2008,26(6)：51-56.
[19]张宾,赵明,董志强,李建国,陈传永,孙锐.作物高产群体lai动态模拟模型的建立与检验[J].作物学报,2007,33(4)：612-619.
[20]周竹青.不同类型小麦品种(系)干物质积累和运转动态比较[J].作物杂志,2002,1：16-19.
[21]高翔,庞红喜,董剑.小麦高产品种光合产物积累与分配及其籽粒灌浆特性研究[J].西北农林科技大学学报(自然科学版),2002,30(4)：24-28.
[22]任正隆,李尧权.小麦开花后的物质积累、籽粒相对生长率和灌浆速率在品种间的变异[J].中国农业科学,1981,14(6)：12-20.
[23]王高武,唐建华,吴维中.小麦抽穗前后干物质生产特点及其对穗重的影响[J].江苏农业学报,1988,4(2)：23-29.
[24]杨连新,王余龙,李世峰,黄建华,董桂春,朱建国,刘钢,韩勇.开放式空气二氧化碳深度增高对小麦物质生产与分配的影响[J].应用生态学报,2007,18(2)：339-346.
[25]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103(1):25-35.
[26]牟会荣,姜东,戴廷波,张传辉,荆奇,曹卫星.遮光对小麦籽粒淀粉品质和花前贮存非结构碳水化合物转运的影响[J].应用生态学报,2009,20(4)：805-810.
[27]王士红.不同年代小麦品种产量与光合特性的演变牲特征[D].南京农业大学.2008
[28]殷美.不同年代小麦品种碳氮物质积累与运转的差异[D].南京农业大学.2008
[29]黄严帅,范袁斌,李炳生,戴学文,张洪程,许轲.氮肥运筹对弱筋小麦宁麦9号群体结构和产量的影响[J].中国农学通报,2008,24(9)：122-126.
[1]Peng S, Cassman K G, Virmani S S, J.Sheehy, Khush G S. Yield potential trends of tropical rice since the release of ir8 and the challenge of increasing rice yield potential[J]. Crop Science,1999,39: 1552-1559.
[2]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[3]李保国,郭焱.作物生长的模拟研究[J].科技导报,1997,7：11-12.
[4]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[5]Peinetti R, Ledent J F. Effect of water status on leaf angle and leaf movement in wheat[J]. Journal of Agronomy and Crop Science,1990,165(4):263-267.
[6]Olesen J E, Berntsen J, Hansen E M, Petersen B M, Petersen J. Crop nitrogen demand and canopy area expansion in winter wheat during vegetative growth[J]. European Journal of Agronomy,2002, 16(4):279-294.
[7]杜永,王艳,王学红,刘辉,杨成,杨建昌.稻麦两熟区超高产小麦株型特征研究[J].麦类作物学报,2008,28(6)：1075-1079.
[8]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[9]胡延吉,兰进好.山东省冬小麦品种冠层结构及光截获的研究[J].中国农业气象,2001,22(3)：28-32.
[10]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[11]陈雨海,余松烈,于振文.小麦生长后期群体光截获量及其分布与产量的关系[J].作物学报,2003,29(5)：730-734.
[12]Aggarwal P K, Kropff M J, Matthews R B, McLaren C G. Using simulation models to design new plant types and to analyse genotype by environment interactions in rice[J]. Plant adaptation and crop improvement,1996:403-418.
[13]吕丽华,赵明,赵久然,陶洪斌,王璞.不同施氮量下夏玉米冠层结构及光合特性的变化不同施氮量下夏玉米冠层结构及光合特性的变化[J].中国农业科学,2008,41(9)：2624-2632.
[14]张强,李自超,吴长明,傅秀林,金京花.不同株穗型水稻超高品种产量构成因素分析[J].西南农业学报,2005,18(5)：518-521.
[15]慕美财,张曰秋,李兴佐,单玉珊.冬小麦高产栽培新途径的研究[J].华北农学报,2008,23(增刊)：167-172.
[2]程式华,曹立勇,庄杰云,吴伟明.关于超级稻品种培育的资源和基因利用问题[J].中国水稻科学,2009,23(3)：223-228.
[3]何中虎,张爱民.中国小麦育种研究进展[M].北京：中国科学技术出版社.2002
[4]魏燮中.我国古农书关于禾谷类作物株型及其生态表现的记载[J].农业考古,1987,1：292-299.
[5]Engledow F L, Wadham S M. Investigations on yield in the cereals[J]. The Journal of Agricultural Science,1923,13:390-439.
[6]陈友订,黄秋妹,张旭.水稻株型育种[M].上海：上海科学技术出版社.2005
[7]Donald C M. The breeding of crop ideotypes[J]. Euphytica,1968,17:385-403.
[8]Matsushima S. Theory and practice of rice cultivation[M]. Tokyo:Yokendo Publishing Co.1968
[9]Mick J J, Pearce R B. An ideotype of maize[J]. Euphytica,1975,24:613-623.
[10]杨守仁,陈温福,张龙步.水稻理想株型育种新动向[J].中国水稻科学,1988,2(3)：129-135.
[11]陈温福,徐正进,张龙步,杨守仁,韩显杰.水稻高产育种研究进展与前景[J].沈阳农业大学学报,1998,2：101-105.
[12]吴永成,周顺利,张永平,王志敏.节水高产小麦理想全株型探讨[J].干旱地区农业研究,2005,23(2)：126-129.
[13]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[14]徐庆章,李登海.玉米株型在高产育种中的作用[J].山东农业科学,1993,3：7-8.
[15]刘建栋,于强,吴乃元.黄淮海地区夏玉米株型结构特征分析[J].中国农业气象,2001,22(1)：6-14.
[16]张其鲁,陈香芝,张立全,李相奎,王家盛.小麦株型分类探讨[J].山东农业科学,2006,1：17-19.
[17]路明,周芳,谢传晓,李明顺,徐云碧,Warburton M,张世煌.玉米杂交种掖单13号的ssr连锁图谱构建与叶夹角和叶向值的qtl定位与分析[J].遗传,2007,29(9)：1131-1138.
[18]裴鑫德.玉米株型的判别松散型[J].华北农学报,1994,9(2)：1-8.
[19]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[20]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[21]郑丕尧.作物生理学导论[M].北京：北京农业大学出版社.1992
[22]Araus J L, Reynolds M P, Acevedo E. Leaf posture, grain yield, growth, leaf structure, and carbon isotope discrimination in wheat[J]. Crop Science,1993,33(6):1273-1279.
[23]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[24]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[25]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[26]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[27]范仲学,王璞,梁振兴,Zilkens M B, Claupein W.优化灌溉与施肥对冬小麦冠层结构的影响研究[J].中国生态农业学报,2005,13(3)：79-81.
[28]Singh P M,张国平,陈锦新.基因型和氮肥对冬小麦叶片形态建成和分蘖的影响[J].麦类作物学报,2000,20(1):28-23. (In English)
[29]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[30]李志勇,郭永新,王璞,翟志席,Zilkens M B.几种水氮模式处理下冬小麦冠层结构的差异[J].麦类作物学报,2007,27(6)：1085-1088.
[31]康祥波,钟阳和,张理,梅楠.冬小麦群体叶层结构的研究[J].河南职业技术师范学院学报,1990,18(3)：13-25.
[32]严威凯.关于小麦株形问题的观察与思考[J].作物杂志,1991,1：14-16.
[33]Annicchiarico P, Pecetti L. Developing a tall durum wheat plant type for semi-arid, mediterranean cereal-livestock farming systems[J]. Field Crops Research,2003,80:157-164.
[34]Marchetti M A. Potential impact of sheath blight on yield and milling quality of short-statured rice lines in the southern united states[J]. Plant Disease,1983,67:162-165.
[35]Sharma N R, Akanda S I, Shahjahan A K M. Development of sheath blight in short, tall, early and late maturing rice cultivars[J]. Bangladesh Journal of Botany,1995,24:143-146.
[36]Han Y P, Xing Y, Gu S, Chen Z, Pan X, Chen X. Effect of morphological traits on sheath blight resistance in rice[J]. Acta Botanica Sinica,2003,45:825-831.
[37]Savary S, Castilla N P, Elazegui F A, McLaren C G, Ynalvez M A, Teng P S. Direct and indirect effects of nitrogen supply and disease source structure on rice sheath blight spread[J]. Phytopathology,1995,85:959-1011.
[38]于经川,刘兆晔,姜鸿明,刘维正.小麦株高、穗幅宽和植株展开度的研究[J].莱阳农学院学报,2001,18(1)：7-11.
[39]张树榛,葛亚新.小麦理想株型研究：小麦株高梯度系列的株型、产量及品质性状的变化研究[J].北京农业大学学报,1990,16(2)：125-132.
[40]雷振生,林作楫.黄淮麦区冬小麦合理株型结构研究[J].华北农学报,1994,4：27-32.
[41]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[42]魏燮中,俞世蓉,宋荷仙,尚勋武.三个生态区小麦株型的比较分析[J].江苏农业学报,1991,7(1)：20-26.
[43]谢令琴,卢少源,吴同彦,王晓燕,常文锁,希慧军.冬小麦早代株高构成指数与产量关系的研究[J].河北农业大学学报,1996,19(1)：12-16.
[44]安呈峰,王延训,毕建杰,叶宝兴.高产小麦生育后期影响茎秆生长的生理因素与抗倒性的关系[J].山东农业科学,2008,7：1-4.
[45]赵万春,王红.小麦株高及其构成因素的遗传和相关性研究[J].麦类作物学报,2003,23(4)：28-31.
[46]吴玉娥,郜庆炉,杨文平,梁云娟,刘俊美.不同小麦品种冠层结构及产量研究[J].河南职业技术师范学院学报,2003,31(3)：1-4.
[47]朱新开,王祥菊,郭凯泉,郭文善,封超年,彭永欣.小麦倒伏的茎秆特征及对产量与品质的影响[J].麦类作物学报,2006,26(1)：87-92.
[48]吴同彦,谢令琴,杨学举,张彩英,陈荣芬.小麦株高构成因素与产量及其他性状相关性的研究[J].河北农业大学学报,2002,25(3)：10-12.
[49]Sharma R C. Tiller mortality and its relationship to grain yield in spring wheat[J]. Field Crops Research,1995,41:55-60.
[50]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var. Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103:25-35.
[51]Jones H G, Kirby E J M. Effects of manipulation of number of tillers and water supply on grain yield in barley[J]. The Journal of Agricultural Science,1977,88:391-397.
[52]Islam T M T, Sedgley R H. Evidence for a "uniculm effect" in spring wheat (triticum aestivum l.) in a mediterranean environment[J]. Euphytica,1981,30:277-282.
[53]李少昆,王崇桃.作物株型和冠层结构信息获取与表达的方法[J].石河子大学学报(自然科学版),1997,1(3)：250-256.
[54]张艳敏,李晋生,钱维朴,黄德明.小麦冠层结构与光分布研究[J].华北农学报,1996,11(1)：54-58.
[55]张立全,张其鲁.小麦株型与田间透光率关系研究[J].潍坊学院学报,2006,6(6)：95-98.
[56]王之杰,郭天财,朱云集,王纪华,赵明.超高产小麦冠层光辐射特征的研究[J].西北植物学报,2003,23(10)：1657-1662.
[57]孙旭生,林琪,李玲燕,姜雯,翟延举.施氮量对超高产小麦生育后期光合特性及产量的影响[J].植物营养与肥料学报,2008,14(5)：840-844.
[58]赵春江,吴华瑞,王纪华,黄文江.田间小麦叶面积空间分布数学模型的建立与应用[J].中国农业科学,2004,37(2)：196-200.
[59]李萌,隋娜,王鹏,赵世杰,孟庆伟,田纪春.生育后期冠层结构对超高产小麦品种(系)光能利用及产量的影响[J].麦类作物学报,2009,29(2)：279-282.
[60]司纪升,王法宏,李升东,冯波,孙令安.不同种植方式对小麦群体质量和产量结构的影响[J].麦类作物学报,2006,26(6)：136-139.
[61]李春明,熊淑萍,赵巧梅,杨颖颖,马新明.有机无机肥配施对小麦冠层结构、产量和蛋白质含量的影响[J].中国农业科学,2008,41(12)：4287-4293.
[62]杨文平,郭天财,刘胜波,王晨阳,王永华,马冬云.行距配置对‘兰考矮早八’小麦后期群体冠层结构及其微环境的影响[J].植物生态学报,2008,32(2)：485-490.
[63]Rasmusson D C. A plant breeder's experience with ideotype breeding[J]. Field Crops Research, 1991,26:191-200.
[64]严威凯.关于小麦株型的观察与思考[J].作物杂志,1991,(1)：14-16.
[65]Yin X, Kropff M J, Goudriaan J, Stam P. A model analysis of yield differences among recombinant inbred lines in barley[J]. Agronomy Journal,2000,92:114-120.
[66]Baker C J, Berry P M, Spink J H, Sylvester-Bradley R, Griffin J M, Scott R K, Clare R W. A method for the assessment of the risk of wheat lodging [J]. Journal of Theoretical Biology,1998, 194(4):587-603.
[67]Berry P M, Sterling M, Baker C J, Spink J H, Sparkes D L. A calibrated model of wheat lodging compared with field measurements[J]. Agricultural and Forest Meteorology,2003,19(4):445-451.
[68]李韬,徐辰武,丁永辉,胡治球,孙长森,顾世梁.一些小麦品种主要株型指标的初步分析[J].扬州大学学报：农业与生命科学版,2003,34(2)：40-43,48.
[69]张其鲁.潍麦8号的株型特征及小麦理想株型研究[D].北京：中国农业大学.2004
[70]吴同彦,谢令琴,葛淑俊,张彩英,王静华,王宝山.小麦营养器官与产量性状相关性的研究[J].河北农业大学学报,2001,24(4)：11-13.
[71]Kropff M J. Application of systems approaches at the field levels[M]. Kluwer Academic Publisher. 1996
[72]Xu Z Z, Yu Z W, Wang D, Zhang Y L. Nitrogen accumulation and translocation for winter wheat under different irrigation regimes[J]. Agronomy and Crop Science,2005,191:439-449.
[73]Ledent J F, Moss D N. Relation of morphological characters and shoot yield in wheat[J]. Crop Science,1979,19(4):445-451.
[74]Yunusa I A M, Siddique K H M, Belford R K, Karimi M M. Effect of canopy structure on efficiency of radiation interception and use in spring wheat cultivars during the pre-anthesis period in a mediterranean-type environment[J]. Field Crops Research,1993,35(2):113-122.
[75]Latiri-Souki K, Nortcliff S, Lawlor D W. Nitrogen fertilizer can increase dry matter, grain production and radiation and water use efficiencies for durum wheat under semi-arid conditions[J]. European Journal of Agronomy,1998,9(1):21-34.
[76]张定一,党建友,王姣爱,裴雪霞,杨武德,苗果园.施氮量对不同品质类型小麦产量、品质和旗叶光合作用的调节效应[J].植物营养与肥料学报,2007,13(4)：535-542.
[77]王月福,姜东,于振文.氮素水平对小麦籽粒产量和蛋白质含量的影响及其生理基础[J].中国农业科学,2003,36(5)：513-520.
[78]周凌云,李卫民.氮素营养对干旱地区小麦光合生理的影响[J].土壤通报,2003,34(3)：195-197.
[79]陆增根,戴廷波,姜东,荆奇,吴正贵,周培南,曹卫星.氮肥运筹对弱筋小麦群体指标与产量和品质形成的影响[J].作物学报,2007,33(4)：590-597.
[80]王月福,于振文,李尚霞,余松烈.土壤肥力和施氮量对小麦根系氮同化及子粒蛋白质含量的影响[J].植物营养与肥料学报,2003,9(1)：39-44.
[81]Pinthus M J. Lodging in wheat, barley, and oats:The phenomenon, its causes, and preventive measures. In Advances in agronomy, vol. Volume 25:Academic Press.209-263.1974.
[82]Kheiralla K A, Mehdi E E, Dawood R A. Evaluation of some wheat cultivars for traits related to lodging resistance under different levels of nitrogen[J]. Assiut Journal of Agricultural Science,1993, 24:257-271.
[83]Crook M J, Ennos A R. The effect of nitrogen and growth regulators on stem and root characteristics associated with lodging in two cultivars of winter wheat[J]. Journal of Experimental Botany,1995,46(8):931-938.
[84]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Growth and morphology of spring wheat (triticum aestivum 1.) culms and their association with lodging: Effects of genotypes, n levels and ethephon[J]. Field Crops Research,2003,84(3):271-290.
[85]张军,戴其根,张洪程,许轲,霍中洋.追氮时期对中筋小麦群体质量及品质的调节效应[J].江苏农业科学,2003,5：34-36.
[86]王月福,于振文,李尚霞,张永丽.不同土壤肥力下强筋小麦适宜施氮量的研究[J].山东农业科学,2001,5：14-15.
[87]Aude B, Christophe L, Christine B, Marie-Helene J. Nitrogen remobilization during grain filling in wheat: Genotypic and environmental effects[J]. Crop Science,2005,45(3):1141.
[88]Arduini I, Masoni A, Ercoli L, Mariotti M. Grain yield, and dry matter and nitrogen accumulation and remobilization in durum wheat as affected by variety and seeding rate[J]. European Journal of Agronomy,2006,25(4):309-318.
[89]Muurinen S, Kleemola J, Peltonen-Sainio P. Accumulation and translocation of nitrogen in spring cereal cultivars differing in nitrogen use efficiency [J]. Agronomy Journal,2007,99(2):441-449.
[90]Ercoli L, Lulli L, Mariotti M, Masoni A, Arduini I. Post-anthesis dry matter and nitrogen dynamics in durum wheat as affected by nitrogen supply and soil water availability[J]. European Journal of Agronomy,2008,28(2):138-147.
[91]王月福,于振文,李尚霞,余松烈.氮素营养水平对小麦开花后碳素同化、运转和产量的影响[J].麦类作物学报,2002,22(2)：55-59.
[92]王月福,于振文,李尚霞,余松烈.土壤肥力和施氮量对小麦氮素吸收运转及籽粒产量和蛋白质含量的影响[J].应用生态学报,2003,14(11)：1868-1872.
[93]沈建辉,戴廷波,荆奇,姜东,曹卫星,郭文善,王龙俊.施氮时期对专用小麦干物质和氮素积累、运转及产量和蛋白质含量的影响[J].作物学报,2004,24(1)：55-58.
[94]吴清丽,高茂盛,廖允成,温晓霞.氮素对冬小麦光合物质贮运及籽粒灌浆进程的影响[J].干旱地区农业研究,2009,27(2)：120-124.
[95]武际,郭熙盛,王允青,汪建来,杨晓虎.氮钾配施对弱筋小麦氮、钾养分吸收利用及产量和品质的影响[J].植物营养与肥料学报,2007,13(6)：1054-1061.
[96]杨延兵,高荣岐,尹燕枰,吴承来,李圣福.不同品质小麦氮素分配及利用率的15n示踪研究[J].麦类作物学报,2008,28(5)：830-835.
[97]同延安,赵营,赵护兵,樊红柱.施氮量对冬小麦氮素吸收、转运及产量的影响[J].植物营养与肥料学报,2007,13(1)：64-69.
[98]石玉,于振文,王东,李延奇,王雪.施氮量和底追比例对小麦氮素吸收转运及产量的影响[J].作物学报,2006,32(12)：1860-1866.
[99]于振文,潘庆民,姜东,张永丽,王东.9000 kg/公顷小麦施氮量与生理特性分析[J].作物学报,2003,29(1)：37-43.
[100]王小燕,于振文.不同施氮量条件下灌溉量对小麦氮素吸收转运和分配的影响[J].中国农业科学,2008,41(10)：3015-3024.
[101]吴海卿,段爱旺,杨传福.冬小麦对不同土壤水分的生理和形态响应[J].华北农学报,2000,5(1)：92-96.
[102]冯广龙,罗远培,杨培岭.节水灌溉对冬小麦干物质分配、灌浆及水分利用率的影响[J].华北农学报,1998,13(2)：11-17.
[103]胡梦芸,张正斌,徐萍,董宝娣,李魏强,李景娟.亏缺灌溉下小麦水分利用效应与光合产 物积累运转的相关研究[J].作物学报,2007,33(10)：1711-1719.
[104]Plaut Z, Butow B J, Blumenthal C S, Wrigley C W. Transport of dry matter into developing wheat kernels and its contribution to grain yield under post-anthesis water deficit and elevated temperature[J]. Field Crops Research,2004,86(2-3):185-198.
[105]王立秋,曹敬山,靳占忠.春小麦产量及品质的水肥效应研究[J].干旱地区农业研究,1997,15(1)：58-63.
[106]王晓燕,于振文.不同施氮量条件下灌溉量对小麦氮素吸收转运和分析的影响[J].中国农业科学,2008,41(10)：3105-3024.
[107]刘萍,郭文善,徐月明,封超年,朱新开,彭永欣.种植密度对中、弱筋小麦籽粒产量和品质的影响[J].麦类作物学报,2006,26(5)：117-121.
[108]吴兰云,周得宝,徐茂林.淮北地区高产小麦的氮肥和密度效应研究[J].中国农学通报,2008,24(7)：153-157.
[109]郭天财,盛坤,冯伟,徐丽娜,王晨阳.种植密度对两种穗型小麦品种分蘖期茎蘖生理特性的影响[J].西北植物学报,2009,29(2)：350-355.
[110]屈会娟,李金才,沈学善,魏凤珍,王成雨,郅胜军.种植密度和播期对冬小麦品种兰考矮早八干物质和氮素积累与转运的影响[J].作物学报,2009,35(1)：124-131.
[111]武兰芳,欧阳竹.不再种植密度下两种穗型小麦叶片光合特性的变化[J].麦类作物学报,2008,28(4)：618-625.
[112]刘万代,杜沛鑫,尹钧,陈现勇.种植密度对超高产小麦豫麦49-198籽粒产量及品质的影响[J].安徽农业科学,2008,36(35)：15402-15404.
[113]于振文,岳寿松,沈成国,余松烈,Egli D B, Sanford V V.不同密度对冬小麦开花后叶片衰老和粒重的影响[J].作物学报,1995,21(4)：412-418.
[114]徐月明.中弱筋小麦优质高产群体质量和株型指标与生理基础研究[D].扬州大学.2004
[115]郭天财,查菲娜,马冬云,宋晓,岳艳军.种植密度对两种穗型冬小麦品种干物质和氮素积累、运转及产量的影响[J].华北农学报,2007,22(6)：152-156.
[116]张晓龙.小麦品种籽粒灌浆研究[J].作物学报,1982,8(2)：87-93.
[117]李金才.品种和播种密度灌浆特性及产量影响的研究[J].安徽农业大学学报,1996,23(4)：461-465.
[118]杨永光,张维成.播量对小麦产量和籽粒营养品质的影响[J].河南职业技术师范学院学报,1989,17(3)：113-116.
[119]董爱民,甘淼,王秀玉,冯新忠,韩林,陈建峰.不同肥力与密度条件下对小麦群体结构及产量的研究[J].河南职业技术师范学院学报,2004,32(2)：12-14.
[1]魏燮中.小麦株型结构分析与产量育种咨询系统[M].南京：东南大学出版社.1991
[2]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[3]张其鲁.潍麦8号的株型特征及小麦理想株型研究[D].北京：中国农业大学.2004
[4]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[5]王士红,荆奇,戴廷波,姜东,曹卫星.不同年代冬小麦品种旗叶光合特性和产量的演变特征[J].应用生态学报,2006,19(6)：1255-1260.
[6]李合生.植物生理生化实验原理和技术[M].北京：高等教育出版社.2000
[7]Gallagher J N. Field studies of cereal leaf growth: Ⅰ. initiation and expansion in relation to temperature and ontogeny[J]. Journal of Experimental Botany,1979,30(4):625-636.
[8]Draper N R, Smith H. Applied regression analysis[M]. John Wiley and Sons.1998
[9]SAS Institute. SAS/STAT 9.1 User's Guide[M]. Cary, NC.2004
[1]任书杰,李世清,王全九,李生秀.栽培模式、施氮和品种对冬小麦冠层结构和产量的影响[J].生态学杂志,2006,25(12)：1449-1454.
[2]Donald C M. The breeding of crop ideotypes[J]. Euphytica,1968,17:385-403.
[3]严威凯.关于小麦株形问题的观察与思考[J].作物杂志,1991,1：14-16.
[4]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[5]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[6]Araus J L, Reynolds M P, Acevedo E. Leaf posture, grain yield, growth, leaf structure, and carbon isotope discrimination in wheat[J]. Crop Science,1993,33(6):1273-1279.
[7]林忠辉,周允华,王辉民,张宜光.青藏高原冬小麦冠层几何结构、光截获及其光合潜能的影响[J].生态学报,1998,18(4)：392-398.
[8]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[9]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[10]Briggs K G, Aytenfisu A. Relationships between morphological characters above the flag leaf node and grain yield in spring wheats[J]. Crop Science,1980,20(3):350-354.
[11]Ledent J F, Moss D M. Relation of morphological characters and shoot yield in wheat[J]. Crop Science,1979,19(4):445-451.
[12]Ishag H M, Mohamed B A, Ishag K H M. Leaf development of spring wheat cultivars in an irrigated heat-stressed environment J]. Field Crops Research,1998,58(3):167-175.
[13]Monneveux P, Reynolds M P, Gonzaez-Santoyo H, Pena R J, Mayr L, Zapata F. Relationships between grain yield, flag leaf morphology, carbon isotope discrimination and ash content in irrigated wheat[J]. Journal of Agronomy and Crop Science,2004,190(6):395-401.
[14]Blake N K, Lanning S P, Martin J M, Sherman J D, Talbert L E. Relationship of flag leaf characteristics to economically important traits in two spring wheat crosses[J]. Crop Science,2007, 47(2):491-494.
[15]王法宏,杨洪宾,徐成忠,李升东,司纪升.垄作栽培对小麦植株形态和产量性状的影响[J].作物学报,2007,33(6)：1038-1040.
[16]Schabenberger O, Pierce F J. Contemporary statistical models for the plant and soil sciences [M]. New York: CRC Press.2002
[17]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[18]陈国庆.小麦形态建成模拟及可视化研究[D].山东农业大学.2004
[19]常丽英.水稻植株形态建成的模拟模型研究[D].南京农业大学.2007
[20]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[21]王义芹,杨兴洪,李滨,童依平,李振声.小麦叶面积及光合速率与产量关系的研究[J].华北农学报,2008,23(增刊)：10-15.
[22]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[23]Singh P M, Zhang G P, Chen J X. Influence of genotypes and nitorgen fertilization on leaf morphogenesis and tillering behaviors in winter wheat[J]. Journal of Triticeae Crops,2000,20(1): 28-23.
[24]李志勇,郭永新,王璞,翟志席,Zilkens M B.几种水氮模式处理下冬小麦冠层结构的差异[J].麦类作物学报,2007,27(6)：1085-1088.
[25]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[26]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[27]伍维模,李世清,王瑞军,邵明安.两种施氮水平下不同基因型冬小麦叶片光合特性与形态差异的研究[J].中国生态农业学报,2006,14(4)：76-81.
[28]Ledent J F. Relationships between grain yield and morphological characters in the winter wheat genotypes of the belgian national list[J]. Biologia Plantarum,1979,21(3):161-169.
[29]Hansen K A, Martin J M, Lanning S P, Talbert L E. Correlation of genotype performance for agronomic and physiological traits in space-planted versus densely seeded conditions[J]. Crop Sci, 2005,45(3):1023-1028.
[30]佟汉文,黄荣华,刘易科,朱展望,张宇庆,秦金成,付汉红,高春保.小麦新品种农艺性状与产量的相关及通径分析[J].湖北农业科学,2008,2008(47)：7.
[31]余泽高,王孝刚.小麦茎叶性状与穗部性状相关性的探讨[J].湖北农学院学报,1998,(1)：8-11.
[32]贾继增.小麦粒重与植株性状相关因素的统计分析[J].作物学报,1984,10(3)：201-205.
[1]魏燮中.小麦植株高度的结构分析[J].南京农业大学学报,1983,1：14-21.
[2]欧阳由男,李春生,章善庆,王会民,朱练峰,禹盛苗,金千瑜,张国平.光周期和有效积温对水稻分蘖角度动态变化的影响[J].应用生态学报,2009,20(5)：1099-1104.
[3]闵东红,翟耀锋,王辉,孟超敏,崔耀锋,李学军,曹宁,于新智.不同株高小麦品种抗倒伏性与其亚性状及产量相关性研究[J].麦类作物学报,2001,21(4)：76-79.
[4]武仙山,王正航,昌小平,景蕊莲.用株高旱胁迫系数分析小麦发育中的抗旱性动态[J].作物学报,2008,34(11)：2010-2018.
[5]赵万春,王红.小麦株高及其构成因素的遗传和相关性研究[J].麦类作物学报,2003,23(4)：28-31.
[6]于经川,刘兆晔,姜鸿明,刘维正,马淑丽.小麦株高整齐度的初步研究[J].莱阳农学院学报,2002,19(1)：20-22.
[7]吴同彦,谢令琴,杨学举,张彩英,陈荣芬.小麦株高构成因素与产量及其他性状相关性的研究[J].河北农业大学学报,2002,25(3)：10-12.
[8]于经川,刘兆晔,姜鸿明,刘维正.小麦株高、穗幅宽和植株展开度的研究[J].莱阳农学院学报,2001,18(1)：7-11.
[9]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var. Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103:25-35.
[10]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Growth and morphology of spring wheat(triticum aestivum 1.) culms and their association with lodging: Effects of genotypes, n levels and ethephon[J]. Field Crops Research,2003,84(3):271-290.
[11]Sharma R C. Tiller mortality and its relationship to grain yield in spring wheat[J]. Field Crops Research,1995,41(1):55-60.
[12]Weiss A, Baenziger P S, McMaster G S, Wilhelm W W, Al Ajlouni Z I. Quantifying phenotypic plasticity using genetic information for simulating plant height in winter wheat[J]. NJAS Wageningen Journal of Life Sciences,2009,57(1):59-64.
[13]白云凤,李希陵.小麦几个数量性状不同生长时期的基因效应分析[J].华北农学报,1997,12(3)：11-16.
[14]蒋向辉,佘朝文,刘伟,何雪梅.青蒿株高生长与分枝特性的曲线方程及其拟合性研究[J].作物杂志,2008,3：46-49.
[15]张殿顺,董翔云,刘树庆.不同施氮水平对春小麦生长发育及其氮素代谢指标的影响[J].华北农学报,2006,21(增刊2)：42-45.
[16]Slafer G A, Halloran G M, Connor D J. Influence of photoperiod on culm length in wheat[J]. Field Crops Research,1995,40(2):95-99.
[17]Tripathi S C, Sayre K D, Kaul J N, Narang R S. Effect of planting methods and n rates on lodging, morphological characters of culm and yield in spring wheat varieties[J]. Cereal Research Communications,2002,30(3-4):431-438.
[18]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[19]傅兆麟.小麦超高产基因型产量因素、株型和冠层结构特征的研究[D].北京：中国农业大学.2001
[20]张娟.超高产小麦品种冠层结构对产量的影响及其遗传研究[D].河南农业大学.2000
[21]胡延吉,兰进好.不同时期小麦主栽品种冠层结构研究[J].中国农业气象,1999,20(1)：11-14.
[22]王淑俭.小麦株高与产量性状相关性的初步研究[J].河南农林科技,1982,7：1-3.
[23]张耀中.小麦株高与产量性状的相关分析[J].河南农业技术师范学院学报,1988,2(1)：59-60.
[24]贾继增.小麦粒重与植株性状相关因素的统计分析[J].作物学报,1984,10(3)：201-205.
[25]庄萍萍,李伟,魏育明,颜泽洪,郑有良.波斯小麦农艺性状相关性及主成分分析[J].麦类作物学报,2006,26(4)：11-14.
[1]胡延吉,兰进好.山东省冬小麦品种冠层结构及光截获的研究[J].中国农业气象,2001,22(3)：28-32.
[2]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[3]吕丽华,赵明,赵久然,陶洪斌,王璞.不同施氮量下夏玉米冠层结构及光合特性的变化不同施氮量下夏玉米冠层结构及光合特性的变化[J].中国农业科学,2008,41(9)：2624-2632.
[4]张玲丽,王辉,孙道杰,冯毅.高产小麦品种冠层形态结构及其产量性状的关系[J].西北植物学报,2004,24(7)：1211-1215.
[5]胡延吉,兰进好,赵坦方,高法振.不同穗型的两个冬小麦品种冠层结构及光合特性的研究[J].作物学报,2000,26(6)：905-912.
[6]赵会杰,李有,邹琦.两个不同穗型小麦品种的冠层辐射和光合特征的比较研究[J].作物学报,2002,28(5)：654-659.
[7]任书杰,李世清,王全九,李生秀.栽培模式、施氮和品种对冬小麦冠层结构和产量的影响[J].生态学杂志,2006,25(12)：1449-1454.
[8]杨文平,郭天财,刘胜波,王晨阳,王永华,马冬云.行距配置对‘兰考矮早八’小麦后期群体冠层结构及其微环境的影响[J].植物生态学报,2008,32(2)：485-490.
[9]熊淑萍,杨颖颖,李春明,马新明,姬兴杰,徐光辉.不同肥料处理对豫麦34冠层结构特征及产量的影响[J].麦类作物学报,2008,28(2)：276-280.
[10]李春明,熊淑萍,赵巧梅,杨颖颖,马新明.有机无机肥配施对小麦冠层结构、产量和蛋白质含量的影响[J].中国农业科学,2008,41(12)：4287-4293.
[11]吴玉娥,郜庆炉,薛香,梁云娟,韩占江.行距对超高产小麦冠层结构及产量构成的影响[J].河南农业科学,2005,9：16-20.
[12]范仲学,王璞,梁振兴,Zilkens M B, Claupein W.优化灌溉与施肥对冬小麦冠层结构的影响研究[J].中国生态农业学报,2005,13(3)：79-81.
[13]张殿顺,董翔云,刘树庆.不同施氮水平对春小麦生长发育及其氮素代谢指标的影响[J].华北农学报,2006,21：42-45.
[14]陆增根,戴廷波,姜东,荆奇,吴正贵,周培南,曹卫星.氮肥运筹对弱筋小麦群体指标与产量和品质形成的影响[J].作物学报,2007,33(4)：590-597.
[15]蔡瑞国,张敏,戴忠民,田雷,王振林.施氮水平对优质小麦旗叶光合特性和子粒生长发育的影响[J].植物营养与肥料学报,2006,12(1)：49-55.
[16]周春菊,张嵩午,王林权,苗芳.施肥对小麦冠层温度的影响及其与生物学性状的关联[J].生 态学报,2005,25(1)：18-22.
[17]孙旭生,林琪,赵长星,翟延举,吴钢,Jaleel C A.施氮量对超高产冬小麦灌浆期旗叶光响应曲线的影响[J].生态学报,2009,29(3)：1428-1437.
[18]赵长星,马东辉,王月福,林琪.施氮量和花后土壤含水量对小麦旗叶衰老及粒重的影响[J].应用生态学报,2008,19(11)：2388-2393.
[19]Hosoi F, Omasa K. Estimating vertical plant area density profile and growth parameters of a wheat canopy at different growth stages using three-dimensional portable lidar imaging[J].ISPRS Journal of Photogrammetry and Remote Sensing,2009,64(2):151-158.
[20]严威凯,王新中.小麦冠层结构中叶面积的层次构成与分析[J].陕西农业科学,1991,3：36-37.
[21]张艳敏,李晋生,钱维朴,黄德明.小麦冠层结构与光分布研究[J].华北农学报,1996,11(1)：54-58.
[22]张娟.超高产小麦品种冠层结构对产量的影响及其遗传研究[D].河南农业大学.2000
[23]Shi Z L, Yin M, Jing Q, Jiang D, Cao W X, Dai T B. Vertical distribution of canopy nitrogen and its relationship with grain protein in winter[J]. Journal of Triticeae Crops,2009,29(2):289-293.
[24]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[25]杨长明,刘敏华,丁超尘,杨林章.三个水稻品种的冠层结构比较研究[J].安徽农业大学学报,2001,28(3)：276-280.
[26]Boedhram N, Arkebauer T J, Batchelor W D. Season-long characterization of vertical distribution of leaf area in corn[J]. Agronomy Journal,2001,93(6):1235-1242.
[27]Subedi K D, Ma B L. Ear position, leaf area, and contribution of individual leaves to grain yield in conventional and leafy maize hybrids[J]. Crop Science,2005,45(6):2246-2257.
[28]王谦,陈景玲,孙治强.LAI-2000冠层分析仪在不同植物群体光分布特征研究中的应用[J].中国农业科学,2006,39(5)：922-927.
[29]章家恩.作物群体建构的生态环境效应及其优化探讨[J].生态科学,2000,19(1)：30-35.
[30]Lotscher M, Stroh K, Schnyder H. Vertical leaf nitrogen distribution in relation to nitrogen status in grassland plants[J]. Annals of Botany,2003,92(5):679-688.
[31]Bertheloot J, Martre P, Andrieu B. Dynamics of light and nitrogen distribution during grain filling within wheat canopy[J]. Plant Physiology,2008,148(3):1707-1720.
[32]Dreccer M F, van Oijen M, Schapendonk A H C M, Pot C S, Rabbinge R. Dynamics of vertical leaf nitrogen distribution in a vegetative wheat canopy: Impact on canopy photosynthesis[J]. Annals of Botany,2000,86(4):821-831.
[33]Kull O. Acclimation of photosynthesis in canopies:Models and limitations[J]. Oecologia,2002, 133(3):267-279.
[34]Pommel B, Gallais A, Coque M, Quiller I, Hirel B, Prioul J L, Andrieu B, Floriot M. Carbon and nitrogen allocation and grain filling in three maize hybrids differing in leaf senescence[J]. European Journal of Agronomy,2006,24(3):203-211.
[35]Field C. Allocating leaf nitrogen for the maximization of carbon gain: Leaf age as a control on the allocation program[J]. Oecologia,1983,56(2):341-347.
[36]Hirose T, Werger M J A. Maximizing daily canopy photosynthesis with respect to the leaf nitrogen allocation pattern in the canopy[J]. Oecologia,1987,72(4):520-526.
[37]王之杰,王纪华,黄文江,马智红,赵明.冬小麦冠层不同叶层和茎鞘氮素与籽粒品质关系的研究[J].中国农业科学,2003,36(12)：1462-1468.
[38]王之杰,王纪华,黄文江,马镥宏,王北洪,赵春江,赵明.冬小麦叶片氮素时空分布特征及其与籽粒品质的关系[J].作物学报,2004,30(7)：700-707.
[39]石祖梁,殷美,荆奇,姜东,曹卫星,戴廷波.冬小麦冠层氮素垂直分布特征及其与籽粒蛋白质的关系[J].麦类作物学报,2009,29(2)：289-293.
[40]秦晓东,戴廷波,荆奇,姜东,曹卫星.冬小麦叶片氮含量时空分布及其与植株氮营养状况的关系[J].作物学报,2006,31(11)：1717-1722.
[41]Milroy S P, Bange M P, Sadras V O. Profiles of leaf nitrogen and light in reproductive canopies of cotton (gossypium hirsutum)[J]. Annals of Botany,2001,87(3):325-333.
[42]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[43]Lv L H, Zhao M, Zhao J R, Tao H B, P W. Canopy structure and photosynthesis of summer maize under different nitrogen fertilizer application rates[J].中国农业科学,2008,41(9):2624-2632.
[1]杨文平.行距和密度对冬小麦冠层结构、微环境及碳氮代谢的影响[D].河南农业大学.2008
[2]姜东,于振文,等.高产冬小麦茎中果聚糖代谢及氮素水平的调控[J].作物学报,2002,28(1)：79-85.
[3]韩占江,郜庆炉,薛香,吴玉娥,敖玲.不同小麦品种干物质积累、分配与产量的关系[J].贵州农业科学,2008,36(6)：14-16.
[4]张利,隋新霞,王羽,樊庆琦,李根英,楚秀生,黄承彦.不同熟性小麦品种的干物质积累和分配规律[J].山东农业科学,2007,6：54-56.
[5]吴清丽,高茂盛,廖允成,温晓霞.氮素对冬小麦光合物质贮运及籽粒灌浆进程的影响[J].干旱地区农业研究,2009,27(2)：120-124.
[6]彭正萍,王艳群,刘淑桥,王红,王蕾,薛世川.不同施肥处理对冬小麦干物质积累及土壤养分垂直分布的影响[J].中国农业科技导报,2007,9(6)：95-99.
[7]马东辉,王月福,周华,孙虎.氮肥和花后土壤含水量对小麦干物质积累、运转及产量的影响[J].麦类作物学报,2007,27(5)：847-851.
[8]屈会娟,李金才,沈学善,魏凤珍,王成雨,郅胜军.种植密度和播期对冬小麦品种兰考矮早八干物质和氮素积累与转运的影响[J].作物学报,2009,35(1)：124-131.
[9]伍维模,李世清,王瑞军,邵明安.两种施氮水平下不同基因型冬小麦叶片光合特性与形态差异的研究[J].中国生态农业学报,2006,14(4)：76-81.
[10]武际,郭熙盛,王允青,汪建来,杨晓虎.氮钾配施对弱筋小麦氮、钾养分吸收利用及产量和品质的影响[J].植物营养与肥料学报,2007,13(6)：1054-1061.
[11]SAS Institute. Sas/stat 9.1 user's guide[M]. Cary, NC.2004
[12]Schabenberger O, Pierce F J. Contemporary statistical models for the plant and soil sciences[M]. Boca Raton:Taylor & Francis/CRC Press.2001
[13]朱庆森,曹显祖,骆亦其.水稻籽粒灌浆的生长分析[J].作物学报,1988,14(3)：182-193.
[14]顾世梁,朱庆森,杨建昌,彭少兵.不同水稻材料籽粒灌浆特性的分析[J].作物学报,2001,27(1)：7-14.
[15]Darroch B A, Baker R J. Grain filling in three spring wheat genotypes:Statistical analysis[J]. Crop Scinece,1990,30:525-529.
[16]沈建辉,戴廷波,荆奇,姜东,曹卫星,郭文善,王龙俊.施氮时期对专用小麦干物质和氮素积累、运转及产量和蛋白质含量的影响[J].麦类作物学报,2004,24(1)：55-58.
[17]王月福,于振文,李尚霞,余松烈.氮素营养水平对小麦开花后碳素同化、运转和产量的影响[J].麦类作物学报,2002,22(2)：55-59.
[18]姚素梅,康跃虎,刘海军.喷灌与地面灌溉冬小麦干物质积累、分配和运转的比较研究[J].干旱地区农业研究,2008,26(6)：51-56.
[19]张宾,赵明,董志强,李建国,陈传永,孙锐.作物高产群体lai动态模拟模型的建立与检验[J].作物学报,2007,33(4)：612-619.
[20]周竹青.不同类型小麦品种(系)干物质积累和运转动态比较[J].作物杂志,2002,1：16-19.
[21]高翔,庞红喜,董剑.小麦高产品种光合产物积累与分配及其籽粒灌浆特性研究[J].西北农林科技大学学报(自然科学版),2002,30(4)：24-28.
[22]任正隆,李尧权.小麦开花后的物质积累、籽粒相对生长率和灌浆速率在品种间的变异[J].中国农业科学,1981,14(6)：12-20.
[23]王高武,唐建华,吴维中.小麦抽穗前后干物质生产特点及其对穗重的影响[J].江苏农业学报,1988,4(2)：23-29.
[24]杨连新,王余龙,李世峰,黄建华,董桂春,朱建国,刘钢,韩勇.开放式空气二氧化碳深度增高对小麦物质生产与分配的影响[J].应用生态学报,2007,18(2)：339-346.
[25]Elhani S, Martos V, Rharrabti Y, Royo C, Garcia del Moral L F. Contribution of main stem and tillers to durum wheat (triticum turgidum 1. Var Durum) grain yield and its components grown in mediterranean environments[J]. Field Crops Research,2007,103(1):25-35.
[26]牟会荣,姜东,戴廷波,张传辉,荆奇,曹卫星.遮光对小麦籽粒淀粉品质和花前贮存非结构碳水化合物转运的影响[J].应用生态学报,2009,20(4)：805-810.
[27]王士红.不同年代小麦品种产量与光合特性的演变牲特征[D].南京农业大学.2008
[28]殷美.不同年代小麦品种碳氮物质积累与运转的差异[D].南京农业大学.2008
[29]黄严帅,范袁斌,李炳生,戴学文,张洪程,许轲.氮肥运筹对弱筋小麦宁麦9号群体结构和产量的影响[J].中国农学通报,2008,24(9)：122-126.
[1]Peng S, Cassman K G, Virmani S S, J.Sheehy, Khush G S. Yield potential trends of tropical rice since the release of ir8 and the challenge of increasing rice yield potential[J]. Crop Science,1999,39: 1552-1559.
[2]封超年,郭文善,何建华,朱新开,马光辉.高产小麦株型的指标体系[J].扬州大学学报自然科学版,1998,1(4)：24-30.
[3]李保国,郭焱.作物生长的模拟研究[J].科技导报,1997,7：11-12.
[4]刘兆晔,于经川,牟春生,姜鸿明,丁晓义.小麦上三叶配置比例对产量性状的影响[J].莱阳农学院学报,2000,17(3)：183-186.
[5]Peinetti R, Ledent J F. Effect of water status on leaf angle and leaf movement in wheat[J]. Journal of Agronomy and Crop Science,1990,165(4):263-267.
[6]Olesen J E, Berntsen J, Hansen E M, Petersen B M, Petersen J. Crop nitrogen demand and canopy area expansion in winter wheat during vegetative growth[J]. European Journal of Agronomy,2002, 16(4):279-294.
[7]杜永,王艳,王学红,刘辉,杨成,杨建昌.稻麦两熟区超高产小麦株型特征研究[J].麦类作物学报,2008,28(6)：1075-1079.
[8]杨文雄.旱地春小麦株型指标与产量形成关系研究[J].干旱地区农业研究,2006,24(1)：43-46.
[9]胡延吉,兰进好.山东省冬小麦品种冠层结构及光截获的研究[J].中国农业气象,2001,22(3)：28-32.
[10]曾浙荣,赵双宋,李青.北京地区高产小麦品种的冠层形成、光截获和产量[J].作物学报,1991,17(3)：161-170.
[11]陈雨海,余松烈,于振文.小麦生长后期群体光截获量及其分布与产量的关系[J].作物学报,2003,29(5)：730-734.
[12]Aggarwal P K, Kropff M J, Matthews R B, McLaren C G. Using simulation models to design new plant types and to analyse genotype by environment interactions in rice[J]. Plant adaptation and crop improvement,1996:403-418.
[13]吕丽华,赵明,赵久然,陶洪斌,王璞.不同施氮量下夏玉米冠层结构及光合特性的变化不同施氮量下夏玉米冠层结构及光合特性的变化[J].中国农业科学,2008,41(9)：2624-2632.
[14]张强,李自超,吴长明,傅秀林,金京花.不同株穗型水稻超高品种产量构成因素分析[J].西南农业学报,2005,18(5)：518-521.
[15]慕美财,张曰秋,李兴佐,单玉珊.冬小麦高产栽培新途径的研究[J].华北农学报,2008,23(增刊)：167-172.