桂林岩溶石山檵木群落不同恢复阶段地上生物量模型构建及分配格局
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摘要
通过AIC、BIC准则结合R~2选择不同恢复阶段的最佳模型,用校正系数估计值的标准误差、平均系统误差和总相对误差值评价所建立模型的精确性,并用所构建的生物量模型对檵木群落不同恢复阶段群落地上生物量和檵木地上生物量的分配格局进行分析。结果表明:(1)运用5种模型进行回归分析,不同恢复阶段树干和地上生物量估测效果极佳,叶和枝生物量次之。选用模型Ⅳ:W=a+b (D~2H)建立乔灌阶段地上生物量的最优模型,选用模型Ⅲ:W=a×D~b×H~c建立灌木阶段和小乔林阶段地上生物量的最优模型。(2)利用建立的檵木群落不同恢复阶段立木生物量生长模型对檵木群落的植被生物量进行估算,不同恢复阶段干生物量和地上生物量大小排列顺序为小乔林阶段>乔灌阶段>灌木阶段;叶生物量和枝生物量大小排列顺序为乔灌阶段>小乔林阶段>灌木阶段。(3)檵木群落中作为建群种的檵木,其地上生物量大小排列顺序为乔灌阶段>小乔林阶段>灌木阶段,檵木生物量在檵木群落不同恢复阶段的群落生物量中所占比例在持续下降。这说明随着桂林岩溶石山檵木群落自然恢复演替的进行,生态系统运行的能量基础和营养物质来源随着群落向更高级的演替阶段发展,而檵木的建群种地位可能会被逐步替代而退居亚乔木层。
The best model was selected by combining AIC and BIC criterions with R~2,simultaneously,the accuracy of the estimation models of above-ground biomass were assessed using the correction coefficient,including standard error of estimated value( SEE),mean systematic error(MSE) and total relative error(TRE),and the distribution pattern of above-ground biomass of Loropetalum chinense and community in different restoration stages of L. chinense communities were analyzed with the established biomass model. The results were as follows:( 1) Five models were used for regression analysis,above-ground biomass and trunk biomass had the best estimate effect,comparing with a lower estimate effect in leaf biomass and branch biomass. We chose Model Ⅳ to establish the optimum model for above-ground biomass of shrub to tree stage,Model Ⅳ was W = a+b( D~2 H). And we chose Model Ⅲ to establish the optimum model for above-ground biomass of shrub stage and smal tree stage,Model Ⅲ was W = a×D~b×H~c.( 2) Based the established optimum model,we estimated that biomass above-ground on the vegetation of L. chinense communities. Among different restoration stages,the leaf biomass and branch biomass order of size was shrub to tree stage > small tree stage > shrub stage,while the aboveground biomass and trunk biomass order of size was small tree stage > shrub to tree stage > shrub stage.( 3) L. chinense as dominant species in L. chinense communities,the order of its above-ground biomass size was shrub to tree stage >small tree stage > shrub stage,and the above-ground biomass of L. chinense contributed to the above-ground biomass of different restoration stages showed a decreasing trend across the succession. This illustrates the energy base and nutrient source of ecosystem operation is progressing toward with the community moves to a higher stage of succession,and the status of edificator of L. chinense in different restoration stages of L. chinense communities may be replaced step by step,and backseat to the sub-tree layer.
The best model was selected by combining AIC and BIC criterions with R~2,simultaneously,the accuracy of the estimation models of above-ground biomass were assessed using the correction coefficient,including standard error of estimated value( SEE),mean systematic error(MSE) and total relative error(TRE),and the distribution pattern of above-ground biomass of Loropetalum chinense and community in different restoration stages of L. chinense communities were analyzed with the established biomass model. The results were as follows:( 1) Five models were used for regression analysis,above-ground biomass and trunk biomass had the best estimate effect,comparing with a lower estimate effect in leaf biomass and branch biomass. We chose Model Ⅳ to establish the optimum model for above-ground biomass of shrub to tree stage,Model Ⅳ was W = a+b( D~2 H). And we chose Model Ⅲ to establish the optimum model for above-ground biomass of shrub stage and smal tree stage,Model Ⅲ was W = a×D~b×H~c.( 2) Based the established optimum model,we estimated that biomass above-ground on the vegetation of L. chinense communities. Among different restoration stages,the leaf biomass and branch biomass order of size was shrub to tree stage > small tree stage > shrub stage,while the aboveground biomass and trunk biomass order of size was small tree stage > shrub to tree stage > shrub stage.( 3) L. chinense as dominant species in L. chinense communities,the order of its above-ground biomass size was shrub to tree stage >small tree stage > shrub stage,and the above-ground biomass of L. chinense contributed to the above-ground biomass of different restoration stages showed a decreasing trend across the succession. This illustrates the energy base and nutrient source of ecosystem operation is progressing toward with the community moves to a higher stage of succession,and the status of edificator of L. chinense in different restoration stages of L. chinense communities may be replaced step by step,and backseat to the sub-tree layer.
引文
BOND,LAMBERTY B,WANG C,2002.Above ground and below ground biomass and sapwood area allometric equations for six boreal tree species of northern Manitoba[J].Can JFor Res,32:1441-1450.
CAI HD,NONG SQ,ZHANG W,et al.,2014.Modeling of standing tree biomass for main species of trees in Guangxi Province[J].For Res Manag,(4):58-61.[蔡会德,农胜奇,张伟,等,2014.广西主要树种立木生物量模型的研建[J].林业资源管理,(4):58-61.]
CHEN PF,LIU CA,ZHANG Y,et al.,2016.Biomass estimating models established for coastal Tamarix chinensis[J].Marine Environ Sci,35(4):551-556.[陈鹏飞,刘长安,张悦,等,2016.滨海湿地柽柳灌丛生物量估算模型[J].海洋环境科学,35(4):551-556.]
CHEN P,ZHOU YC,2017.Soil nutrient capacity and forest tree sustainability in plateau karst region[J].Earth Environ,45(1):32-37.[陈萍,周运超,2017.高原喀斯特土壤养分容量及其林木持续性研究[J].地球与环境,45(1):32-37.]
CHEN Y,FENG T,YU YL,et al.,2017.The litter dynamics of three plant communities in karst forest of western Guizhou Province[J].J Guizhou Univ Eng Sci,35(3):73-78.[陈元,冯图,喻元良,等,2017.黔西北喀斯特山区三种植物群落凋落物动态研究[J].贵州工程应用技术学院学报,35(3):73-78.]
CUI LL,TU YL,LA D,et al.,2017.Establishment of biomass prediction model for six typical shrubs in Lasa River Basin[J].Tibet Sci Tech,7:64-68.[崔玲玲,土艳丽,拉多,等,2017.拉萨河流域6种典型灌木生物量预测模型的建立[J].西藏科技,7:64-68.]
HUANG CD,ZHANG J,YANG WQ,et al.,2008.Dynamics on forest carbon stock in Sichuan Province and Chongqing City[J].Acta Ecol Sin,28(3):966-975.[黄从德,张健,杨万勤,等,2008.四川省及重庆地区森林植被碳储量动态[J].生态学报,28(3):966-975.]
LI G,ZHAO X,ZHANG BB,et al.,2014.Biomass allocation pattern and Caragana korshinskii with estimation model building of different plant heights[J].Acta Agr Sin,22(4):769-775.[李刚,赵祥,张宾宾,等,2014.不同株高的柠条生物量分配格局及其估测模型构建[J].草地学报,22(4):769-775.]
LI HK,ZHAO PX,LEI YC,et al.,2012.Comparison on estimation of wood biomass using forest inventory data[J].Sci Silv Sin,48(5):44-52.[李海奎,赵鹏祥,雷渊才,等,2012.基于森林清查资料的乔木林生物量估算方法的比较[J].林业科学,48(5):44-52.]
LI L,XIA FC,SUN Y,et al.,2017.Biomass allocation of eight early-spring herbs in broadleaf Pinus koraiensis mixed forest[J].J Beijing For Univ,39(1):34-42.[李良,夏富才,孙越,等,2017.阔叶红松林下早春植物生物量分配[J].北京林业大学学报,39(1):34-42.]
LI SD,HU SP,TANG XM,2013.Dynamic change of carbon storage in forest vegetation[M].Beijing:Science Press.[李世东,胡淑萍,唐小明,2013.森林植被碳储量动态变化研究[M].北京:科学出版社.]
LI W,WANG CK,ZHANG QZ,2015.Differentiation of stand individuals impacts allometry and biomass allocation of Larix gmelinii trees[J].Acta Ecol Sin,35(6):1679-1687.[李巍,王传宽,张全智,2015.林木分化对兴安落叶松异速生长方程和生物量分配的影响[J].生态学报,35(6):1679-1687.]
LI XK,HE CX,TANG JS,et al.,2008.Evolution and ecological processes of karst ecosystem of Guangxi[J].Guangxi Sci,15(1):80-86,91.[李先琨,何成新,唐建生,等,2008.广西岩溶山地生态系统特征与恢复重建[J].广西科学,15(1):80-86,91.]
LI Z,ZHAO YJ,SONG HY,et al.,2017.The effects of soil thickness heterogeneity on grassland plant community structure and growth of dominant species in karst area[J].Pratac Sci,34(10):2023-2032.[李周,赵雅洁,宋海燕,等,2017.喀斯特土层厚度异质性对草地群落结构和优势种生长的影响[J].草业科学,34(10):2023-2032.]
LIU QJ,2009.Nested regression for establishing tree biomass equations[J].Chin J Plant Ecol,33(2):331-337.[刘琪璟,2009.嵌套式回归建立树木生物量模型[J].植物生态学报,33(2):331-337.]
LIU YS,DENG XS,HU YC,2006.Rocky land degradation and poverty alleviation strategy in Guangxi karst mountainous area[J].Mt Res,24(4):228-233.[刘彦随,邓旭升,胡业翠,2006.广西喀斯特山区土地石漠化与扶贫开发探析[J].山地学报,24(4):228-233.]
LUO QB,ZENG WS,HE DB,et al.,1999.Establishment and application of compatible tree above-ground biomass models[J].J Nat Resour,14(3):271-277.[骆期邦,曾伟生,贺东北,等,1999.立木地上部分生物量模型的建立及其应用研究[J].自然资源学报,14(3):271-277.]
MA JM,LIANG SC,LIANG MY,et al.,2009.Aboveground biomass and its allocation of main shrub types in karst hills of Guilin,China[J].J Guangxi Norm Univ(Nat Sci Ed),27(4):95-98.[马姜明,梁士楚,梁月明,等,2009.桂林岩溶石山主要灌丛类型地上生物量及分配特征[J].广西师范大学学报(自然科学版),27(4):95-98.]
MA JM,WU M,ZHAN TT,et al.,2013.Changes of species composition and diversity among restoration stages of Loropetalum chinense communities in karst area of Lijiang River valley[J].Ecol Environ Sci,22(1):66-71.[马姜明,吴蒙,占婷婷,等,2013.漓江流域岩溶区檵木群落不同恢复阶段物种组成及多样性变化[J].生态环境学报,22(1):66-71.]
MA JM,ZHAN TT,MO ZY,et al.,2012.Changes of niche among restoration stages of Loropetalum chinense communities in karst area of Lijiang River Valley[J].Acta Bot BorealOccident Sin,32(12):2530-2536.[马姜明,占婷婷,莫祖英,等,2012.漓江流域岩溶区檵木群落不同恢复阶段主要共有种生态位变化[J].西北植物学报,32(12):2530-2536.]
MEI JL,ZHUANG FH,MA JM,et al.,2017.Spatial point pattern analysis of Alchornea trewioides population clonal growth in the karst area of Guilin[J].Acta Ecol Sin,37(9):3164-3171.[梅军林,庄枫红,马姜明,等,2017.桂林喀斯特地区克隆生长红背山麻杆种群的点格局分析[J].生态学报,37(9):3164-3171.]
MONIKA K,TIBOR SK,KLAUDIA K,2012.Comparison of soil microbial communities from two distinct karst areas in Hungary[J].Acta Microbiol Imm H,59(1):91-105.
ODUM EP,BARRETT GW,1971.Fundamentals of ecology[M].Philadelphia:Saunders.
QIN YH,MA JM,MEI JL,et al.,2017.The initial dynamic of litter decomposition of Loropetalum chinense communities among different recovery stages in karst area of Lijiang River watershed[J].Acta Ecol Sin,20:6792-6799.[覃扬浍,马姜明,梅军林,等,2017.漓江流域岩溶区檵木群落不同恢复阶段凋落物分解初期动态[J].生态学报,20:6792-6799.]
SU RL,2017.Allometric equations to estimate biomass of different age-sequence of Chinese fir(Cunninghamia lanceolata)plantations[J].J Fujian For Sci Technol,44(2):105-108.[苏瑞兰,2017.异速生长模型估算不同林龄杉木人工林生物量[J].福建林业科技,44(2):105-108.]
WANG JF,OU GL,TANG JR,et al.,2012.Biomass estimation model of shrub community at Jatropha curcas growing area in Lincang of Yunnan[J].J W Chin For Sci,41(6):53-57.[王俊峰,欧光龙,唐军荣,等,2012.临沧膏桐种植区灌木群落生物量估测模型研究[J].西部林业科学,41(6):53-57.]
WANG XK,FENG ZW,OUYANG ZY,2001.Vegetation carbon storage and density of forest ecosystems in China[J].Chin J Appl Ecol,12(1):13-16.[王效科,冯宗炜,欧阳志云,2001.中国森林生态系统的植物碳储量和碳密度研究[J].应用生态学报,12(1):13-16.]
WANG ZC,DU H,SONG TQ,et al.,2015.Allometric models of major tree species and forest biomass in Guangxi[J].Acta Ecol Sin,35(13):4462-4472.[汪珍川,杜虎,宋同清,等,2015.广西主要树种(组)异速生长模型及森林生物量特征[J].生态学报,35(13):4462-4472.]
WANG WF,LEI YC,WANG XF,et al.,2008.A review of forest biomass models[J].J NW For Univ,23(2):58-63.[王维枫,雷渊才,王雪峰,等,2008.森林生物量模型综述[J].西北林学院学报,23(2):58-63.]
WANG Y,XU WT,XIONG GM,et al.,2017.Biomass allocation patterns of Loropetalum chinense[J].Chin J Plant Ecol,41(1):105-114.[王杨,徐文婷,熊高明,等,2017.檵木生物量分配特征[J].植物生态学报,41(1):105-114.]
WEI NF,2017.Comparative analysis and establishment of biomass for Pinus elliottii of different altitudes in Northern Fujian[J].For Prosp Des,(2):61-64.[魏年锋,2017.闽北不同海拔湿地松生物量模型构建及比较分析[J].林业勘察设计,(2):61-64.]
YANG XL,WEI XR,SHAO MA,2016.Stem biomass estimation models for dominant shrubs on the northern Loess Plateau of China[J].Chin J Appl Ecol,27(10):3164-3172.[杨宪龙,魏孝荣,邵明安,2016.黄土高原北部典型灌丛枝条生物量估算模型[J].应用生态学报,27(10):3164-3172.]
ZENG WS,TANG SZ,2011.Goodness evaluation and precision analysis of tree biomass equations[J].Sci Silv Sin,47(11):106-113.[曾伟生,唐守正,2011.立木生物量方程的优度评价和精度分析[J].林业科学,47(11):106-113.]
ZENG WS,XIAO QH,HU J,et al.,2010.Establishment of single tree biomass equations for Pinus massoniana in southern China[J].J Centr S Univ For Technol,30(5):50-56.[曾伟生,肖前辉,胡觉,等,2010.中国南方马尾松立木生物量模型研建[J].中南林业科技大学学报,30(5):50-56.]
ZENG WS,2014.Comparison of three allometric equations for biomass modeling[J].Centr S For Invent Plan,33(1):1-3.[曾伟生,2014.3种异速生长方程对生物量建模的对比分析[J].中南林业调查规划,33(1):1-3.]
ZHU J,HAN HR,KANG FF,et al.,2016.Biomass allocation patterns and allometric models of Larix principis-rupprechtii in Mt.Taiyue,Shanxi[J].J Ecol,35(11):2918-2925.[朱江,韩海荣,康峰峰,等,2016.山西太岳山华北落叶松生物量分配格局与异速生长模型[J].生态学杂志,35(11):2918-2925.]
ZHU SQ,WEI LM,CHEN ZR,et al.,1995.A preliminary study on biomass components of karst forest in Maolan of Guizhou Province,China[J].Acta Phytoecol Sin,19(4):358-367.[朱守谦,魏鲁明,陈正仁,等,1995.茂兰喀斯特森林生物量构成初步研究[J].植物生态学报,19(4):358-367.]
CAI HD,NONG SQ,ZHANG W,et al.,2014.Modeling of standing tree biomass for main species of trees in Guangxi Province[J].For Res Manag,(4):58-61.[蔡会德,农胜奇,张伟,等,2014.广西主要树种立木生物量模型的研建[J].林业资源管理,(4):58-61.]
CHEN PF,LIU CA,ZHANG Y,et al.,2016.Biomass estimating models established for coastal Tamarix chinensis[J].Marine Environ Sci,35(4):551-556.[陈鹏飞,刘长安,张悦,等,2016.滨海湿地柽柳灌丛生物量估算模型[J].海洋环境科学,35(4):551-556.]
CHEN P,ZHOU YC,2017.Soil nutrient capacity and forest tree sustainability in plateau karst region[J].Earth Environ,45(1):32-37.[陈萍,周运超,2017.高原喀斯特土壤养分容量及其林木持续性研究[J].地球与环境,45(1):32-37.]
CHEN Y,FENG T,YU YL,et al.,2017.The litter dynamics of three plant communities in karst forest of western Guizhou Province[J].J Guizhou Univ Eng Sci,35(3):73-78.[陈元,冯图,喻元良,等,2017.黔西北喀斯特山区三种植物群落凋落物动态研究[J].贵州工程应用技术学院学报,35(3):73-78.]
CUI LL,TU YL,LA D,et al.,2017.Establishment of biomass prediction model for six typical shrubs in Lasa River Basin[J].Tibet Sci Tech,7:64-68.[崔玲玲,土艳丽,拉多,等,2017.拉萨河流域6种典型灌木生物量预测模型的建立[J].西藏科技,7:64-68.]
HUANG CD,ZHANG J,YANG WQ,et al.,2008.Dynamics on forest carbon stock in Sichuan Province and Chongqing City[J].Acta Ecol Sin,28(3):966-975.[黄从德,张健,杨万勤,等,2008.四川省及重庆地区森林植被碳储量动态[J].生态学报,28(3):966-975.]
LI G,ZHAO X,ZHANG BB,et al.,2014.Biomass allocation pattern and Caragana korshinskii with estimation model building of different plant heights[J].Acta Agr Sin,22(4):769-775.[李刚,赵祥,张宾宾,等,2014.不同株高的柠条生物量分配格局及其估测模型构建[J].草地学报,22(4):769-775.]
LI HK,ZHAO PX,LEI YC,et al.,2012.Comparison on estimation of wood biomass using forest inventory data[J].Sci Silv Sin,48(5):44-52.[李海奎,赵鹏祥,雷渊才,等,2012.基于森林清查资料的乔木林生物量估算方法的比较[J].林业科学,48(5):44-52.]
LI L,XIA FC,SUN Y,et al.,2017.Biomass allocation of eight early-spring herbs in broadleaf Pinus koraiensis mixed forest[J].J Beijing For Univ,39(1):34-42.[李良,夏富才,孙越,等,2017.阔叶红松林下早春植物生物量分配[J].北京林业大学学报,39(1):34-42.]
LI SD,HU SP,TANG XM,2013.Dynamic change of carbon storage in forest vegetation[M].Beijing:Science Press.[李世东,胡淑萍,唐小明,2013.森林植被碳储量动态变化研究[M].北京:科学出版社.]
LI W,WANG CK,ZHANG QZ,2015.Differentiation of stand individuals impacts allometry and biomass allocation of Larix gmelinii trees[J].Acta Ecol Sin,35(6):1679-1687.[李巍,王传宽,张全智,2015.林木分化对兴安落叶松异速生长方程和生物量分配的影响[J].生态学报,35(6):1679-1687.]
LI XK,HE CX,TANG JS,et al.,2008.Evolution and ecological processes of karst ecosystem of Guangxi[J].Guangxi Sci,15(1):80-86,91.[李先琨,何成新,唐建生,等,2008.广西岩溶山地生态系统特征与恢复重建[J].广西科学,15(1):80-86,91.]
LI Z,ZHAO YJ,SONG HY,et al.,2017.The effects of soil thickness heterogeneity on grassland plant community structure and growth of dominant species in karst area[J].Pratac Sci,34(10):2023-2032.[李周,赵雅洁,宋海燕,等,2017.喀斯特土层厚度异质性对草地群落结构和优势种生长的影响[J].草业科学,34(10):2023-2032.]
LIU QJ,2009.Nested regression for establishing tree biomass equations[J].Chin J Plant Ecol,33(2):331-337.[刘琪璟,2009.嵌套式回归建立树木生物量模型[J].植物生态学报,33(2):331-337.]
LIU YS,DENG XS,HU YC,2006.Rocky land degradation and poverty alleviation strategy in Guangxi karst mountainous area[J].Mt Res,24(4):228-233.[刘彦随,邓旭升,胡业翠,2006.广西喀斯特山区土地石漠化与扶贫开发探析[J].山地学报,24(4):228-233.]
LUO QB,ZENG WS,HE DB,et al.,1999.Establishment and application of compatible tree above-ground biomass models[J].J Nat Resour,14(3):271-277.[骆期邦,曾伟生,贺东北,等,1999.立木地上部分生物量模型的建立及其应用研究[J].自然资源学报,14(3):271-277.]
MA JM,LIANG SC,LIANG MY,et al.,2009.Aboveground biomass and its allocation of main shrub types in karst hills of Guilin,China[J].J Guangxi Norm Univ(Nat Sci Ed),27(4):95-98.[马姜明,梁士楚,梁月明,等,2009.桂林岩溶石山主要灌丛类型地上生物量及分配特征[J].广西师范大学学报(自然科学版),27(4):95-98.]
MA JM,WU M,ZHAN TT,et al.,2013.Changes of species composition and diversity among restoration stages of Loropetalum chinense communities in karst area of Lijiang River valley[J].Ecol Environ Sci,22(1):66-71.[马姜明,吴蒙,占婷婷,等,2013.漓江流域岩溶区檵木群落不同恢复阶段物种组成及多样性变化[J].生态环境学报,22(1):66-71.]
MA JM,ZHAN TT,MO ZY,et al.,2012.Changes of niche among restoration stages of Loropetalum chinense communities in karst area of Lijiang River Valley[J].Acta Bot BorealOccident Sin,32(12):2530-2536.[马姜明,占婷婷,莫祖英,等,2012.漓江流域岩溶区檵木群落不同恢复阶段主要共有种生态位变化[J].西北植物学报,32(12):2530-2536.]
MEI JL,ZHUANG FH,MA JM,et al.,2017.Spatial point pattern analysis of Alchornea trewioides population clonal growth in the karst area of Guilin[J].Acta Ecol Sin,37(9):3164-3171.[梅军林,庄枫红,马姜明,等,2017.桂林喀斯特地区克隆生长红背山麻杆种群的点格局分析[J].生态学报,37(9):3164-3171.]
MONIKA K,TIBOR SK,KLAUDIA K,2012.Comparison of soil microbial communities from two distinct karst areas in Hungary[J].Acta Microbiol Imm H,59(1):91-105.
ODUM EP,BARRETT GW,1971.Fundamentals of ecology[M].Philadelphia:Saunders.
QIN YH,MA JM,MEI JL,et al.,2017.The initial dynamic of litter decomposition of Loropetalum chinense communities among different recovery stages in karst area of Lijiang River watershed[J].Acta Ecol Sin,20:6792-6799.[覃扬浍,马姜明,梅军林,等,2017.漓江流域岩溶区檵木群落不同恢复阶段凋落物分解初期动态[J].生态学报,20:6792-6799.]
SU RL,2017.Allometric equations to estimate biomass of different age-sequence of Chinese fir(Cunninghamia lanceolata)plantations[J].J Fujian For Sci Technol,44(2):105-108.[苏瑞兰,2017.异速生长模型估算不同林龄杉木人工林生物量[J].福建林业科技,44(2):105-108.]
WANG JF,OU GL,TANG JR,et al.,2012.Biomass estimation model of shrub community at Jatropha curcas growing area in Lincang of Yunnan[J].J W Chin For Sci,41(6):53-57.[王俊峰,欧光龙,唐军荣,等,2012.临沧膏桐种植区灌木群落生物量估测模型研究[J].西部林业科学,41(6):53-57.]
WANG XK,FENG ZW,OUYANG ZY,2001.Vegetation carbon storage and density of forest ecosystems in China[J].Chin J Appl Ecol,12(1):13-16.[王效科,冯宗炜,欧阳志云,2001.中国森林生态系统的植物碳储量和碳密度研究[J].应用生态学报,12(1):13-16.]
WANG ZC,DU H,SONG TQ,et al.,2015.Allometric models of major tree species and forest biomass in Guangxi[J].Acta Ecol Sin,35(13):4462-4472.[汪珍川,杜虎,宋同清,等,2015.广西主要树种(组)异速生长模型及森林生物量特征[J].生态学报,35(13):4462-4472.]
WANG WF,LEI YC,WANG XF,et al.,2008.A review of forest biomass models[J].J NW For Univ,23(2):58-63.[王维枫,雷渊才,王雪峰,等,2008.森林生物量模型综述[J].西北林学院学报,23(2):58-63.]
WANG Y,XU WT,XIONG GM,et al.,2017.Biomass allocation patterns of Loropetalum chinense[J].Chin J Plant Ecol,41(1):105-114.[王杨,徐文婷,熊高明,等,2017.檵木生物量分配特征[J].植物生态学报,41(1):105-114.]
WEI NF,2017.Comparative analysis and establishment of biomass for Pinus elliottii of different altitudes in Northern Fujian[J].For Prosp Des,(2):61-64.[魏年锋,2017.闽北不同海拔湿地松生物量模型构建及比较分析[J].林业勘察设计,(2):61-64.]
YANG XL,WEI XR,SHAO MA,2016.Stem biomass estimation models for dominant shrubs on the northern Loess Plateau of China[J].Chin J Appl Ecol,27(10):3164-3172.[杨宪龙,魏孝荣,邵明安,2016.黄土高原北部典型灌丛枝条生物量估算模型[J].应用生态学报,27(10):3164-3172.]
ZENG WS,TANG SZ,2011.Goodness evaluation and precision analysis of tree biomass equations[J].Sci Silv Sin,47(11):106-113.[曾伟生,唐守正,2011.立木生物量方程的优度评价和精度分析[J].林业科学,47(11):106-113.]
ZENG WS,XIAO QH,HU J,et al.,2010.Establishment of single tree biomass equations for Pinus massoniana in southern China[J].J Centr S Univ For Technol,30(5):50-56.[曾伟生,肖前辉,胡觉,等,2010.中国南方马尾松立木生物量模型研建[J].中南林业科技大学学报,30(5):50-56.]
ZENG WS,2014.Comparison of three allometric equations for biomass modeling[J].Centr S For Invent Plan,33(1):1-3.[曾伟生,2014.3种异速生长方程对生物量建模的对比分析[J].中南林业调查规划,33(1):1-3.]
ZHU J,HAN HR,KANG FF,et al.,2016.Biomass allocation patterns and allometric models of Larix principis-rupprechtii in Mt.Taiyue,Shanxi[J].J Ecol,35(11):2918-2925.[朱江,韩海荣,康峰峰,等,2016.山西太岳山华北落叶松生物量分配格局与异速生长模型[J].生态学杂志,35(11):2918-2925.]
ZHU SQ,WEI LM,CHEN ZR,et al.,1995.A preliminary study on biomass components of karst forest in Maolan of Guizhou Province,China[J].Acta Phytoecol Sin,19(4):358-367.[朱守谦,魏鲁明,陈正仁,等,1995.茂兰喀斯特森林生物量构成初步研究[J].植物生态学报,19(4):358-367.]