After the revision of all the known species of the genus Leucocythere, occurred in the Late Cenozoic strata from the Qaidam Basin (Qinghai), their common character has been found: there are developed and often furcated anterior cardinal tooth on the right valves. It differs from the diagnosis of the genus Leucocythere. Among these species, two form of the intercardinal bar are present, one of them with crenulated or notched intercardinal bar, and the another one with the intercardinal bar really smooth. Paleoenvironmental distribution of Denticulocythere and Potamocythere in Eurasia suggests that most Denticulocythere are brackish water species, while Potamocythere live in the fresh or brackish water.